Badamia exclamationis
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''Badamia exclamationis'', commonly known as the brown awl or narrow-winged awl,Markku Savela's website on Lepidoptera. Page on genu

TOL web page on genu
''Badamia''
/ref> is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group compris ...
belonging to the family
Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...
. It is found in
south South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...
and
southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
,
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
, and
Oceania Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
.


Range

The brown awl is found in
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand ...
, South
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...
,
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
and
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
. This butterfly is found throughout the
Indian subcontinent The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, In ...
and in the
Andaman islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between t ...
. The type locality is
South India South India, also known as Dakshina Bharata or Peninsular India, consists of the peninsular southern part of India. It encompasses the Indian states of Andhra Pradesh, Karnataka, Kerala, Tamil Nadu, and Telangana, as well as the union terr ...
.


Status

As per William Harry Evans (1932), the butterfly is common in India and rare in the
Andaman islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between t ...
.
Mark Alexander Wynter-Blyth Mark Alexander Wynter-Blyth (15 August 1906 – 16 April 1963 Leysin, Switzerland) was an English schoolteacher and amateur naturalist who wrote one of the first field guides to the butterflies of the Indian region. He was also involved in censuses ...
(1957) records it as "Not Rare" and "Locally Common". Krushnamegh Kunte (2000) reports it as common in deciduous forests during the monsoon months and the evergreen forests in the following months.


Habits

A forest butterfly, the brown awl favours openings and edges of deciduous and evergreen forests while its caterpillars are to be found in moist deciduous and semi-evergreen forests. It flies about either late or early in the morning in the shade of the jungles. It can be sometimes seen in bright sunlight visiting flowers, such as ''
Glycosmis ''Glycosmis'' is a genus of flowering plants in the citrus family, Rutaceae and tribe Clauseneae.''Glyc ...
'', ''
Buddleia ''Buddleja'' (; ''Buddleia''; also historically given as ''Buddlea'') is a genus comprising over 140 species of flowering plants endemic to Asia, Africa, and the Americas. The generic name bestowed by Linnaeus posthumously honoured the Revere ...
'', ''
Chromolaena ''Chromolaena'' is a genus of about 165 species of perennials and shrubs in the family Asteraceae. The name is derived from the Greek words (), meaning "color", and () or () meaning "cloak". It refers to the colored phyllaries of some species. ...
'' and ''
Lantana ''Lantana'' () is a genus of about 150 species of perennial flowering plants in the verbena family, Verbenaceae. They are native to tropical regions of the Americas and Africa but exist as an introduced species in numerous areas, especially in ...
'', but is very wary and energetic at such times, moving jerkily and rapidly between flowers or across
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
s. It can also be seen
mud-puddling Mud-puddling, or simply puddling, is a behaviour most conspicuous in butterflies, but occurs in other animals as well, mainly insects; they seek out nutrients in certain moist substances such as rotting plant matter, mud and carrion and they suck ...
or at bird droppings. During a population explosion, like those of the common banded awl (''
Hasora chromus ''Hasora chromus'', the common banded awl,Markku Savela's website on Lepidoptera – page on genu is a butterfly belonging to the family Hesperiidae which is found in the Indian subcontinent, Southeast Asia and Australia. Range The common ban ...
''), the caterpillars of the brown awl may strip away all their food supply forcing the butterflies to migrate to other places where a fresh supply of host plants is available and even to other habitats such as shrubs, grasslands and gardens. The brown awl flies as low as 6 feet over the bushes or as high as 60 to 75 feet in the canopy. The adults feed at lower levels on flowers of shrubs and small trees, but ascend to higher reaches of the vegetation to lay eggs or to bask, which it does very occasionally, holding its wings flat with the forewings covering the hindwings thus giving an
arrowhead An arrowhead or point is the usually sharpened and hardened tip of an arrow, which contributes a majority of the projectile mass and is responsible for impacting and penetrating a target, as well as to fulfill some special purposes such as sign ...
effect. The flight of the butterfly is fast and bounding with an audible wing beat. When inactive, it rests on the undersides of leaves in shady forest spots, with the head pointing downwards. If disturbed it will generally buzz around energetically before returning to the same spot to rest.


Description

The brown awl is a non-descript brown butterfly, darker above and lighter below. The sexes are alike, except for three to four semi-transparent spots on the forewing which cannot be differentiated in the field. The skipper has a light-brown abdomen with black bands across it. The dry-season form is usually smaller, paler, and may not have the forewing spots. This skipper is unmistakable because of its long and narrow wings. It has the longest wings in proportion to breadth of all Indian butterflies.


Detailed description

Edward Yerbury Watson (1891) gives a detailed description: Genus characters Species description


Similar species

Unlike the other awls, the brown awl lacks the narrow white wing bands on the hindwings. The very distinctive characteristics of the brown awl are the characteristic shape of the body and the narrower wings than the other awls.


Life cycle


Eggs

The brown awl lays many eggs on a single plant, one at a time, on the tips of fresh shoots. The dome-shaped egg is pale green with longitudinal ridges having fine beadings; a total 13 ridges in all.


Larva

The larva is a pale violaceous (violet) yellow, with numerous black transverse dorsal lines; the prolegs are whitish encircled with black. The head is yellow, approximately heart shaped, with a black band and many tiny black spots. On hatching the larva webs the edges of leaves together with silk to form a roomy cell from a leaf in which it resides throughout the larval stage. When disturbed, it can move quite briskly and even drop off. The caterpillars of the brown awl grow faster than most of those of other families, and have moist, sticky droppings. At the time of pupation they descend close to the ground, looking for suitable spots to pupate. The caterpillar constructs a tubular cell from a leaf by drawing the edges together with thick strands of silk. In this cell, the caterpillar prepares an extensive silk bed on which it sits awaiting pupation. The freshly formed pupa clings onto the silken pad almost immediately.


Pupa

The pupa is stubby, with protruding eyes and a prominent projection on the head in between them. The pupa may be light brown or violaceous. The body tapers away from the shoulders towards the rear. The abdomen is creamish with a row of four black spots on each side. The pupa is shiny, but plastered with a white powder.


Host plants

The caterpillars have been recorded on the following deciduous and semi-evergreen forest plants, mostly from the family
Combretaceae The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lianas in ca 10 genera. The family includes the leadwood tree, ''Combret ...
: * Large climbing shrubs of ''
Combretum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but th ...
'' such as '' Combretum albidum'', '' Combretum latifolium'' and ''
Combretum ovalifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but ...
''. * Large forest trees of genus '' Terminalia'' such as '' Terminalia bellerica'', ''
Terminalia catappa ''Terminalia catappa'' is a large tropical tree in the leadwood tree family, Combretaceae, native to Asia, Australia, the Pacific, Madagascar and Seychelles. Common names in English include country almond, Indian almond, Malabar almond, sea almon ...
'' and '' Terminalia oblongata''. * ''
Anogeissus acuminata ''Terminalia phillyreifolia'' is an Asian species of tree in the family Combretaceae. It has been called buttontree or yon (from ; IPA: ). It is a medium-sized tree found in both primary and secondary tropical and sub-tropical forests. It is rec ...
'' and '' Chionanthus purpureus''. * ''
Hiptage benghalensis ''Hiptage benghalensis'', often simply called hiptage, is a perennial plant, perennial, evergreen liana native to India, Southeast Asia, Taiwan, and the Philippines. Its habitat is variable and prefers climates ranging from warm temperate to trop ...
'' (
Malpighiaceae Malpighiaceae is a family of flowering plants in the order Malpighiales. It comprises about 73 genera and 1315 species, all of which are native to the tropics and subtropics. About 80% of the genera and 90% of the species occur in the New Wor ...
)nhm.ac.uk Caterpillar Host plant database
/ref> * '' Linociera purpurea''. * ''
Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extendi ...
'' spp.


Gallery

image:Badamia exclamationis (ento-csiro-au).jpg, Brown awl Image:ExclamationisCRW 5480-01.jpg, Museum specimens File:7淡綠弄蝶5(劉威良攝) (31621207561).jpg, Dorsal view File:7淡綠弄蝶21(李榮芳攝) (30927073453).jpg, Mating pair


See also

* Coeliadinae * List of butterflies of India (Coeliadinae) *
List of butterflies of India (Hesperiidae) India has a rich biodiversity of butterflies, of which skippers are a well represented family. Of the seven subfamilies belonging to the family Hesperiidae, four are found in India, comprising a total of 223 species of 74 genera and these are li ...


References


Bibliography

* * Brower, Andrew V. Z. and Warren, Andrew, (2007). Coeliadinae Evans 1937. Version 21 February 2007 (temporary). http://tolweb.org/Coeliadinae/12150/2007.02.21 in The Tree of Life Web Project, http://tolweb.org/ * * * * Watson, E. Y. (1891) ''Hesperiidae indicae''. Vest and Co. Madras. *


External links


Images representing ''Badamia exclamationis''
at
Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...
{{DEFAULTSORT:Badamia Exclamationis Coeliadinae Butterflies of Asia Butterflies of Oceania Butterflies of Indochina Butterflies of Malaysia Butterflies of Singapore Butterflies described in 1775 Taxa named by Johan Christian Fabricius