''Austriadraco'' is a genus of

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...

living during the

Late Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch ...

in the area of present

Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...

. Its only species—''Austriadraco dallavecchiai''—was previously attributed to ''Eudimorphodon'', and its closest relatives may have been ''

Eudimorphodon ''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (m ...

'' or ''

Arcticodactylus ''Arcticodactylus'' is a genus of basal pterosaur living during the Late Triassic in the area of present Greenland. Its only species was previously attributed to ''Eudimorphodon'', and its closest relatives may have been ''Eudimorphodon'' or ''A ...

''.

Discovery

In June 1994, near Seefeld in Austrian

Tirol Tyrol (; historically the Tyrole; de-AT, Tirol ; it, Tirolo) is a historical region in the Alps - in Northern Italy and western Austria. The area was historically the core of the County of Tyrol, part of the Holy Roman Empire, Austrian Emp ...

, at a 1600 metres high mountain trail to the ''

Reither Spitze The Reither Spitze is a mountain in the Karwendel in the Austrian federal state of Tyrol that, thanks to its location at the southwestern end of the Erlspitze Group, is particularly well known for its great views of the surrounding area from its ...

'', in the vicinity of the ''Reither Joch-Alm'', Bernd Lammerer discovered a pterosaur skeleton. The remains have been secured as five stone plates, removed on several occasions. In 2003,

Peter Wellnhofer Peter Wellnhofer (born Munich, 1936) is a German paleontologist at the Bayerische Staatssammlung fur Paläontologie in Munich. He is best known for his work on the various fossil specimens of ''Archaeopteryx'' or "Urvogel", the first known bird. W ...

identified the fossil as a specimen of ''

'', a cf. ''E. ranzii''. As it was 10 to 25% shorter than the latter's holotype, Wellnhofer considered it a juvenile. The same year

Fabio Marco Dalla Vecchia Fabio is a given name descended from Latin '' Fabius'' and very popular in Italy and Latin America (due to Italian migration). Its English equivalent is Fabian. The name is written without an accent in Italian and Spanish, but is usually accented ...

doubted the comparability to ''E. ranzii'' and suggested that it represent a separate ''Eudimorphodon'' species. In 2009, Dalla Vecchia concluded that the specimen was neither a juvenile nor closely related to ''Eudimorphodon''. In 2015,

Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...

named the separate genus ''Austriadraco'', with the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...

''Austriadraco dallavecchiai''. The generic name is a combination of the

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

words ''Austria'' and ''draco'', "dragon". The specific name honours Dalla Vecchia. The

Life Science Identifiers Life Science Identifiers are a way to name and locate pieces of information on the web. Essentially, an LSID is a unique identifier for some data, and the LSID protocol specifies a standard way to locate the data (as well as a standard way of descr ...

are for the genus 120B3003-6DE3-41B4-AF6B-6F242FB2A777 and for the species 6E123721-07EA-419CB755-9981CC7D9209. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

, BSP 1994 I 51, was found in a layer of the Seefeld Formation, dating from the late

Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...

. It consists of a partial and disarticulated skeleton with skull. It contains both frontal bones, a left jugal, the lower jaws, loose teeth, vertebrae of the neck, back and tail, the shoulder girdle, both humeri, a first wing phalanx, the pelvis, a shinbone and a calf bone. The fused frontals had in 2003 been incorrectly identified as a breast bone by Wellnhofer. The bones have been partly preserved as impressions only and many are fragmented. The fossil is part of the collection of the '' Bayerische Staatssammlung für Paläontologie und historische Geologie'' at

Munich Munich ( ; german: München ; bar, Minga ) is the capital and most populous city of the German state of Bavaria. With a population of 1,558,395 inhabitants as of 31 July 2020, it is the third-largest city in Germany, after Berlin and ...

Description

''Austriadraco dallavecchiai'' is a small species.

Humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a r ...

length is about four centimetres. In 2015 Kellner indicated several distinguishing traits. Some of these are

autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...

. The

frontal bone The frontal bone is a bone in the human skull. The bone consists of two portions.'' Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, ...

has a short front branch. The

jugal bone The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Ana ...

has short branches to the front, in the direction of the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...

and the

nasal bone The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Ea ...

, and a long narrow upwards branch running towards the

postorbital bone The ''postorbital'' is one of the bones in vertebrate skulls which forms a portion of the dermal skull roof and, sometimes, a ring about the orbit. Generally, it is located behind the postfrontal and posteriorly to the orbital fenestra. In some v ...

. In the outer rear side of the lower jaw an opening is present, the mandibular fenestra. The coronoid process of the

surangular The suprangular or surangular is a jaw bone found in most land vertebrates, except mammals. Usually in the back of the jaw, on the upper edge, it is connected to all other jaw bones: dentary, angular, splenial and articular. It is often a mu ...

bone is low. The shoulder blade is considerably longer, 62%, than the

coracoid A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is prese ...

. Additionally, a unique combination of in themselves not unique traits is present. The coracoid is broad, with a constricted shaft. In the pelvis, the ischipubic plate, the fusion of the

pubic bone In vertebrates, the pubic region ( la, pubis) is the most forward-facing ( ventral and anterior) of the three main regions making up the coxal bone. The left and right pubic regions are each made up of three sections, a superior ramus, inferior ...

with the

ischium The ischium () form ...

, is deep. The shinbone is relatively long, with a length of 57.7 millimetres attaining 70% of the length of the humerus and 92% of the length of the first phalanx of the (fourth) wingfinger.

Classification

According to Dalla Vecchia's analysis, ''Austriadraco'' would have a very basal position in the Pterosauria. Kellner concluded that its affinities were uncertain and placed ''Austriadraco'' in a separate, undefined,

Austriadraconidae Caviramidae is a group of basal pterosaurs. It was erected by paleontologist Matthew G. Baron in 2020. It was defined as the least inclusive clade that includes ''Arcticodactylus cromptonellus'' and ''Caviramus schesaplanensis''. The members of ...

. He suggested a close relationship with ''

'' as both taxa shared the trait of a short coracoid. In 2020, a study upheld by Matthew G. Baron about early pterosaur interrelationships found ''Austriadraco'' to group with ''

Carniadactylus ''Carniadactylus'' is a genus of pterosaur which existed in Europe during the Late Triassic period (late Carnian or early Norian, about 228 million years agoBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribu ...

'', ''

Raeticodactylus ''Raeticodactylus'' is a genus of non-pterodactyloid pterosaur from the late Norian-early Rhaetian-age Upper Triassic lower Kössen Formation of the central Austroalpine of Grisons, Switzerland. It is known from holotype BNM 14524, a single dis ...

'', and the Austriadraconidae, which in turn were within a clade called

Caviramidae Caviramidae is a group of basal pterosaurs. It was erected by paleontologist Matthew G. Baron in 2020. It was defined as the least inclusive clade that includes ''Arcticodactylus cromptonellus'' and '' Caviramus schesaplanensis''. The members o ...

.Matthew G. Baron (2020). "Testing pterosaur ingroup relationships through broader sampling of avemetatarsalian taxa and characters and a range of phylogenetic analysis techniques". PeerJ. 8: e9604. doi:10.7717/peerj.9604. PMC 7512134. .

References

{{Taxonbar, from=Q20652509 Pterosaurs Norian life Late Triassic pterosaurs of Europe Triassic Austria Fossil taxa described in 2015 Taxa named by Alexander Kellner

Discovery

Description

Classification

See also

References