Austriadactylus
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''Austriadactylus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of "
rhamphorhynchoid The Rhamphorhynchoidea forms one of the two suborders of pterosaurs and represents an evolutionary grade of primitive members of flying reptiles. This suborder is paraphyletic unlike the Pterodactyloidea, which arose from within the Rhamphorhync ...
"
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
. The fossil remains were unearthed in
Late Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch ...
(middle
Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...
ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107

/ref>) rocks of
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
. The genus was named in 2002 by
Fabio Marco Dalla Vecchia Fabio is a given name descended from Latin '' Fabius'' and very popular in Italy and Latin America (due to Italian migration). Its English equivalent is Fabian. The name is written without an accent in Italian and Spanish, but is usually accented ...
e.a.. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
is ''Austriadactylus cristatus''. The genus name is derived from
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
''Austria'' and Greek ''daktylos'', "finger", in reference to the wing finger of pterosaurs. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
means "crested" in Latin, a reference to the skull crest. The genus is based on
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
SMNS 56342, a crushed partial skeleton on a slab, found in an abandoned mine near Ankerschlag in
Tyrol Tyrol (; historically the Tyrole; de-AT, Tirol ; it, Tirolo) is a historical region in the Alps - in Northern Italy and western Austria. The area was historically the core of the County of Tyrol, part of the Holy Roman Empire, Austrian Emp ...
, in the
Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...
Seefelder Beds. The counterslab has been lost and with it some of the bone. The fossil consists of the skull, lower jaws, some
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
e, parts of the limbs and pelvic girdle, and the first part of the tail. The elongated skull has a length of 11 cm. It carried a bony crest that widened as it descended towards the snout, up to height of 2 cm. The triangular nares formed the largest skull openings. The also triangular ''fenestrae antorbitales'' are smaller than the orbits. The teeth differ in shape and the species was thus heterodont. Most teeth are small and tricuspid or three-pointed. In the front of the upper jaw five larger recurved teeth with a single point form a prey grab; six or seven such teeth are also interspersed with the smaller teeth more to the back of the mouth. There are at least seventeen and perhaps as much as 25 tricuspid teeth in the upper jaw, for a total of perhaps 74 teeth of all sizes in the skull. The number of teeth in the lower jaws cannot be determined. The flexible tail did not have the stiffening rod-like vertebral extensions present in other basal pterosaurs. The wingspan has been estimated at about 120 cm. ''Austriadactylus'' was in 2002 assigned by the describers to a general Pterosauria ''
incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'', but some later analyses showed it to have been related to '' Campylognathoides'' and ''
Eudimorphodon ''Eudimorphodon'' was a pterosaur that was discovered in 1973 by Mario Pandolfi in the town of Cene, Italy and described the same year by Rocco Zambelli. The nearly complete skeleton was retrieved from shale deposited during the Late Triassic (m ...
'' in the
Campylognathoididae Novialoidea (meaning "new wings") is an extinct clade of macronychopteran pterosaurs that lived from the latest Early Jurassic to the latest Late Cretaceous (early Toarcian to late Maastrichtian ageBarrett, P. M., Butler, R. J., Edwards, N. P ...
. It has even been suggested it was a junior synonym of ''Eudimorphodon'', though perhaps a distinct species in that genus. The following phylogenetic analysis follows the topology of Upchurch ''et al.'' (2015).


See also

* Timeline of pterosaur research


References

*


External links


The Pterosaur Database (pdf)
{{Taxonbar, from1=Q19298186, from2=Q2789205 Pterosaurs Late Triassic pterosaurs of Europe Fossil taxa described in 2002 Fossils of Austria