Augacephalus
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''Augacephalus'' is a genus of harpacterine theraphosid
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s. It has three species, all of which are found in Africa.


Taxonomy and etymology

The type species of ''Augacephalus'' is ''A. breyeri'' which was described as '' Pterinochilus breyeri'' by Hewitt in 1919. In 2002, Gallon placed it a new genus, which he erected in the same paper, ''Augacephalus''.Gallon, R. C. (2002). Revision of the African genera ''Pterinochilus'' and ''Eucratoscelus'' (Araneae, Theraphosidae, Harpactirinae) with description of two new genera. ''Bulletin of the British Arachnological Society'' 12: 201-232 Its name comes from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
αυγή ''auga'' meaning "sun rays" and κεφᾰλή ''kephale'' meaning "head" which refers to the prominent, radial cephalothorax
striae Stretch marks, also known as striae () or striae distensae, are a form of scarring on the skin with an off-color hue. Over time they may diminish, but will not disappear completely. Striae are caused by tearing of the dermis during periods of r ...
present in most species.


Natural history

All known species are fossorial and females lay eggs in a hammock egg-sack which yield about 95
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
lings.Gallon, R. C. (2010). On some southern African Harpactirinae, with notes on the eumenophorines ''Pelinobius muticus'' Karsch, 1885 and ''Monocentropella'' Strand, 1907 (Araneae, Theraphosidae). ''Bulletin of the British Arachnological Society'' 15: 29-48


Diagnosis

''Augacephalus'' is separated from other harpactirines in the following ways: Distinguished from '' Harpactirella'' by the presence of a retrolateral
chelicera The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or similarly ...
l
scopula ''Scopula'' is a genus of moths in the family Geometridae described by Franz von Paula Schrank in 1802. Species It has 705 species which are listed here alphabetically. A *'' Scopula ablativa'' (Dognin, 1911) *''Scopula abolita'' Herbulot, ...
composed of plumose
seta In biology, setae (singular seta ; from the Latin word for " bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. ...
e (in males scopula not obviously composed of plumose setae). Separated from '' Harpactira'' and '' Trichognatha'' by the absence of a dense scopula on the upper prolateral cheliceral surface. Further separated from ''Harpactira'' by the absence of plumose stridulatory strikers on the prolateral maxillary surface, and by the absence of a discrete row of bristles below the retrolateral cheliceral scopula. Distinguished from '' Idiothele'' by the possession of digitiform distal segment on posterior
spinneret A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and a ...
s. Differs from '' Ceratogyrus'' by the lack of a
foveal The fovea centralis is a small, central pit composed of closely packed cones in the eye. It is located in the center of the macula lutea of the retina. The fovea is responsible for sharp central vision (also called foveal vision), which is ...
tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection ...
/procurved fovea. Female ''Augacephalus'' are separated from those of '' Eucratoscelus'' by the unmodified (not incrassate) tibiae of leg IV. Male ''Augacephalus'' are separated from those of ''Eucratoscelus'' and '' Pterinochilus'' by lacking the distal proventral tibial apophysis or by the reduced surmounted megaspine. Further separated from ''Eucratoscelus'' by the absence of a distal proventral tumid protuberance on
metatarsus The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the me ...
I. The presence of a distal prodorsal spine on metatarsi III and IV further separates ''Augacephalus'' from ''Eucratoscelus''. Female ''Augacephalus'' are separated from those of ''Pterinochilus'' by the absence of long emergent setae on the chelicerae (giving them a velvety appearance), their robust
palp Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") ...
i and legs I–II, and by the position of their posterior
sternal The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Sh ...
sigilla (an impressed sclerotized spot).


Species

The
World Spider Catalog The World Spider Catalog (WSC) is an online searchable database concerned with spider taxonomy. It aims to list all accepted families, genera and species, as well as provide access to the related taxonomic literature. The WSC began as a series of ...
accepts the following species: *'' Augacephalus breyeri'' Hewitt, 1919 South Africa, Mozambique, Eswatini (
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
) *'' Augacephalus ezendami'' Gallon, 2001 Mozambique *'' Augacephalus junodi'' Simon, 1904 East, South Africa


References

{{Taxonbar, from=Q2608814 Theraphosidae Theraphosidae genera Spiders of Africa