The Atlantic halibut (''Hippoglossus hippoglossus'') is a

flatfish A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating ...

of the family

Pleuronectidae Pleuronectidae, also known as righteye flounders, are a family of flounders. They are called "righteye flounders" because most species lie on the sea bottom on their left sides, with both eyes on their right sides. The Paralichthyidae are the op ...

. They are

demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...

living on or near sand, gravel or clay bottoms at depths of between . The halibut is among the largest

teleost Teleostei (; Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all extant species of fish. Tele ...

(bony) fish in the world, and is a

threatened species Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of '' critical depen ...

due to a slow rate of growth and

overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in t ...

. Halibut are strong swimmers and are able to migrate long distances. Halibut size is not age-specific, but rather tends to follow a cycle related to halibut (and therefore food) abundance. The native habitat of the Atlantic halibut is the temperate and arctic waters of the northern

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

, from

Labrador , nickname = "The Big Land" , etymology = , subdivision_type = Country , subdivision_name = Canada , subdivision_type1 = Province , subdivision_name1 ...

and

Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...

Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...

, the

Barents Sea The Barents Sea ( , also ; no, Barentshavet, ; russian: Баренцево море, Barentsevo More) is a marginal sea of the Arctic Ocean, located off the northern coasts of Norway and Russia and divided between Norwegian and Russian terr ...

and as far south as the

Bay of Biscay The Bay of Biscay (), known in Spain as the Gulf of Biscay ( es, Golfo de Vizcaya, eu, Bizkaiko Golkoa), and in France and some border regions as the Gulf of Gascony (french: Golfe de Gascogne, oc, Golf de Gasconha, br, Pleg-mor Gwaskogn), ...

and

Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth are ...

. It is the largest flatfish in the world, reaching lengths of up to and weights of . Its lifespan can reach 50 years. Age can be estimated by counting the rings laid down inside the otolith – a bony structure found inside the inner ear of the fish.

Description

The Atlantic halibut is a right-eyed flounder. It is flattened sideways and habitually lies on the left side of its body with both eyes migrating to the right side of its head during development. When the larvae are born, they start in the upright position like most other fish having one eye on each side of the head. Once the larvae reach one inch in length, the left eye moves over the snout to the right side of the head. At the same time, the coloring on the left side begins to fade to white while the upper side of the fish is a uniformly dark chocolate, olive or slate color, and can be almost black. The end of the caudal fin is concave. Young fish are paler with more mottled coloration.Atlantic halibut
Gulf of Maine Research Institute: ''Fishery Bulletin'' (February, 2006) Male adults average about but can range to upwards of , rarely reaching . Females can get as large as .

Biology

Atlantic Halibut have a relatively slow growth rate and late onset of sexual maturity, with males attaining maturity at seven to eight years old, females at 10 to 11 years, and individuals averagely live around 25 to 30 years old. The oldest to have been caught was 50 years old. The Alaskan Halibut is a batch spawner where females lay anywhere from a few thousand to four million eggs (depending on the size and age of the fish, older females tend to lay only two million in one spawning period). Spawning occurs between December to April near the bottom of the ocean between . The egg size is around 3.0–3.8 mm and the larvae at time of hatching is 6.5 mm. Growth weight is dependent on the density, competition, and availability of food.

Habitat

This marine fish usually lives on the ocean floor at depths between , but it occasionally comes closer to the surface. The larvae are

pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...

, drifting relatively helplessly, but at around 4 cm, they migrate to the bottom. Young between the ages of two and four years live close to the shore, moving into deeper waters as they grow older.

Geographic distribution

Found in both the eastern and western portions of the North Atlantic. In the western Atlantic, found from southwestern Greenland and Labrador, Canada to Virginia in the USA. They are found in the eastern Atlantic around Iceland, the United Kingdom, and northern Europe to Russia. A map of the Atlantic Halibut's geographic distribution can be found on the Official Website of Massachusetts Division of Marine Fisheries. This demersal fish is found ranging from the latitude and longitude coordinates: 79°N - 36°N, 77°W - 55°E.

Role in ecosystem

The Atlantic halibut occupies a relatively high

trophic level The trophic level of an organism is the position it occupies in a food web. A food chain is a succession of organisms that eat other organisms and may, in turn, be eaten themselves. The trophic level of an organism is the number of steps it ...

in the

food chain A food chain is a linear network of links in a food web starting from producer organisms (such as grass or algae which produce their own food via photosynthesis) and ending at an apex predator species (like grizzly bears or killer whales), de ...

Diet

The diet of the Atlantic halibut consists mainly of other fish, e.g. cod, haddock,

herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Ocean ...

, pogge,

sand eel Sand eel or sandeel is the common name used for a considerable number of species of fish. While they are not true eels, they are eel-like in their appearance and can grow up to in length. Many species are found off the western coasts of Europe ...

s and

capelin The capelin or caplin (''Mallotus villosus'') is a small forage fish of the smelt family found in the North Atlantic, North Pacific and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capelin ...

, but it will also eat

cephalopod A cephalopod is any member of the molluscan class Cephalopoda ( Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head ...

s, large

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...

s and other

benthos Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone.

Predators

Atlantic halibut are eaten by

seal Seal may refer to any of the following: Common uses * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to imp ...

s, and are a

staple food A staple food, food staple, or simply a staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard Diet (nutrition), diet for a given person or group of people, supplying a large fraction of ...

of the

Greenland shark The Greenland shark (''Somniosus microcephalus''), also known as the gurry shark, grey shark, or by the Kalaallisut name ''eqalussuaq'', is a large shark of the family Somniosidae ("sleeper sharks"), closely related to the Pacific and souther ...

. Killer whales are also natural predators of the halibut. Humans also consume halibut.

Commercial fishing of wild Atlantic halibut

The wild Atlantic halibut was formerly a very important food fish, but due to its slow rate of population growth, it is unable to recover quickly from overfishing, and the fishery has largely collapsed. Consequently, wild fish labelled as "halibut" are usually one of the other large flatfishes, including

Pacific halibut The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

, ''Hippoglossus stenolepis''. The most popular way to catch the Alaskan halibut is to use circle hooks that are baited with herring or other similar fish, fished from the bottom with cannonball weights of up to 36 ounces on a slider. The survival rate of caught and released halibut is 95 percent due to the lack of having a swim bladder and not severely suffering from changes in water pressure (as long as the captured halibut is in excellent condition).

Farming

Due to its popularity as a food fish, Atlantic halibut has attracted investment in

fish farming upright=1.3, Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye">mariculture.html" ;"title="Salmon farming in the sea (mariculture">Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye, Scotland Fish farming o ...

. As of 2006, five countries—Canada, Norway, the UK, Iceland and Chile—were engaged in some form of Atlantic halibut

aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...

production.

Conservation status

In 1996, the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

rated it as

Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...

and placed it on its

Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...

. The Atlantic halibut is a US

National Marine Fisheries Service The National Marine Fisheries Service (NMFS), informally known as NOAA Fisheries, is a United States federal agency within the U.S. Department of Commerce's National Oceanic and Atmospheric Administration (NOAA) that is responsible for the stew ...

Species of Concern, one of those species about which the U.S. Government's

National Oceanic and Atmospheric Administration The National Oceanic and Atmospheric Administration (abbreviated as NOAA ) is an United States scientific and regulatory agency within the United States Department of Commerce that forecasts weather, monitors oceanic and atmospheric conditi ...

has some concerns regarding status and threats, but for which insufficient information is available to indicate a need to list the species under the U.S.

Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...

(ESA). The American Fisheries Society has classified the species as "Vulnerable". In 2010, Greenpeace International added the Atlantic halibut to its seafood red list of "fish that are commonly sold in supermarkets around the world, and which have a very high risk of being sourced from unsustainable fisheries."Greenpeace International Seafood Red list
The Atlantic halibut has been on the endangered species list since 1996 due to overfishing since it became commercially important during the 19th century since the great meat demand of the halibut and being one of the largest flatfishes and essentially being easy to capture.

References

External links

Video of larval Atlantic halibut reared in a tank and feeding on Artemia
{{Taxonbar, from=Q1143520

Atlantic halibut The Atlantic halibut (''Hippoglossus hippoglossus'') is a flatfish of the family Pleuronectidae. They are demersal fish living on or near sand, gravel or clay bottoms at depths of between . The halibut is among the largest teleost (bony) fish in ...

Fish of the North Sea Fish of the North Atlantic Fauna of Atlantic Canada Fauna of the British Isles