Asiatic lion
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The Asiatic lion is a population of '' Panthera leo leo'' that today survives in the wild only in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
. Since the turn of the 20th century, its range has been restricted to Gir National Park and the surrounding areas in the Indian state of
Gujarat Gujarat (, ) is a state along the western coast of India. Its coastline of about is the longest in the country, most of which lies on the Kathiawar peninsula. Gujarat is the fifth-largest Indian state by area, covering some ; and the ninth ...
. Historically, it inhabited much of southwest Asia to northern India. The first
scientific description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have b ...
of the Asiatic lion was published in 1826 by the
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
n zoologist Johann N. Meyer, who named it ''Felis leo persicus''. On the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
, it is listed under its former
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''Panthera leo persica'' as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
because of its small population size and
area of occupancy Area is the quantity that expresses the extent of a region on the plane or on a curved surface. The area of a plane region or ''plane area'' refers to the area of a shape or planar lamina, while ''surface area'' refers to the area of an open su ...
. Until the 19th century, it occurred in
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the Ara ...
, eastern
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
,
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Mesopotamia Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the ...
,
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
, and from east of the
Indus River The Indus ( ) is a transboundary river of Asia and a trans-Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in Western Tibet, flows northwest through the disputed region of Kashmi ...
to
Bengal Bengal ( ; bn, বাংলা/বঙ্গ, translit=Bānglā/Bôngô, ) is a geopolitical, cultural and historical region in South Asia, specifically in the eastern part of the Indian subcontinent at the apex of the Bay of Bengal, predom ...
and the Narmada River in
Central India Central India is a loosely defined geographical region of India. There is no clear official definition and various ones may be used. One common definition consists of the states of Chhattisgarh and Madhya Pradesh, which are included in al ...
. The population has steadily increased since 2010. In May 2015, the 14th Asiatic Lion Census was conducted over an area of about ; the lion population was estimated at 523 individuals, comprising 109 adult males, 201 adult females and 213 cubs. In August 2017, surveyors counted 650 wild lions. In June 2020, an estimation exercise counted 674 Asiatic lions in the Gir forest region, an increase of 29% over the 2015 census figure. The lion is one of five pantherine cats native to India, along with the
Bengal tiger The Bengal tiger is a population of the '' Panthera tigris tigris'' subspecies. It ranks among the biggest wild cats alive today. It is considered to belong to the world's charismatic megafauna. The tiger is estimated to have been present i ...
(''P. tigris tigris''), Indian leopard (''P. pardus fusca''),
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a felid in the genus '' Panthera'' native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is es ...
(''P. uncia'') and
clouded leopard The clouded leopard (''Neofelis nebulosa''), also called the mainland clouded leopard, is a wild cat inhabiting dense forests from the foothills of the Himalayas through mainland Southeast Asia into South China. In the early 19th century, a ...
(''Neofelis nebulosa''). It was also known as the Indian lion and the Persian lion.


Taxonomy

''Felis leo persicus'' was the scientific name proposed by Johann N. Meyer in 1826 who described an Asiatic lion skin from
Persia Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
. In the 19th century, several zoologists described lion
zoological specimen A zoological specimen is an animal or part of an animal preserved for scientific use. Various uses are: to verify the identity of a (species), to allow study, increase public knowledge of zoology. Zoological specimens are extremely diverse. Exampl ...
from other parts of Asia that used to be considered
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are al ...
s of ''P. l. persica'': *''Felis leo bengalensis'' proposed by
Edward Turner Bennett Edward Turner Bennett (6 January 1797 – 21 August 1836) was an English zoologist and writer. He was the elder brother of the botanist John Joseph Bennett.
in 1829 was a lion kept in the
menagerie A menagerie is a collection of captive animals, frequently exotic, kept for display; or the place where such a collection is kept, a precursor to the modern zoological garden. The term was first used in 17th-century France, in reference to the m ...
of the
Tower of London The Tower of London, officially His Majesty's Royal Palace and Fortress of the Tower of London, is a historic castle on the north bank of the River Thames in central London. It lies within the London Borough of Tower Hamlets, which is sep ...
. Bennett's essay contains a drawing titled 'Bengal lion'. *''Felis leo goojratensis'' proposed by Walter Smee in 1833 was based on two skins of maneless lions from Gujarat that Smee exhibited in a meeting of the
Zoological Society of London The Zoological Society of London (ZSL) is a charity devoted to the worldwide conservation of animals and their habitats. It was founded in 1826. Since 1828, it has maintained the London Zoo, and since 1931 Whipsnade Park. History On 29 ...
. *''Leo asiaticus'' proposed by
Sir William Jardine, 7th Baronet Sir William Jardine, 7th Baronet of Applegarth FRS FRSE FLS FSA (23 February 1800 – 21 November 1874) was a Scottish naturalist. He is known for his editing of a long series of natural history books, ''The Naturalist's Library''. Life an ...
in 1834 was a lion from India. *''Felis leo indicus'' proposed by Henri Marie Ducrotay de Blainville in 1843 was based on an Asiatic lion
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
. In 2017, the Asiatic lion was subsumed to ''P. l. leo'' due to close morphological and
molecular genetic Molecular genetics is a sub-field of biology that addresses how differences in the structures or expression of DNA molecules manifests as variation among organisms. Molecular genetics often applies an "investigative approach" to determine the ...
similarities with Barbary lion specimens. However, several scientists continue using ''P. l. persica'' for the Asiatic lion.


Evolution

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
remains of '' Panthera spelaea'' excavated in the Cromer Stage indicate that it represented a genetically isolated and highly distinct lineage, not closely related to Asiatic lions. Fossil lion remains were found in
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
deposits in
West Bengal West Bengal (, Bengali: ''Poshchim Bongo'', , abbr. WB) is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of . West Bengal is the fou ...
. A fossil carnassial excavated in the Batadomba Cave indicates that the Sri Lanka lion (''P. l. sinhaleyus'') inhabited
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
during the
late Pleistocene The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch withi ...
, and is thought to have become extinct around 39,000 years ago.
Deraniyagala Deraniyagala is a town in the Kegalle district in the Sabaragamuwa Province of Sri Lanka. Deraniyaga city area consists major government administrative offices, textile industries and transportation hub and the district hospital which has seve ...
described this lion in 1939.


Phylogeography

Results of a phylogeographic analysis based on
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
sequences of lions from across the global range, including now extinct populations like Barbary lions, indicates that
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
n lions are
phylogenetically In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...
basal to all modern lions. These findings support an African origin of modern lion evolution with a probable centre in East and
Southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
. It is likely that lions migrated from there to West Africa, eastern
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
and via the periphery of the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...
into Turkey,
southern Europe Southern Europe is the southern region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countries and regions: Alb ...
and northern India during the last 20,000 years. The
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
, Congolian rainforests and the Great Rift Valley are natural barriers to lion dispersal.
Genetic marker A genetic marker is a gene or DNA sequence with a known location on a chromosome that can be used to identify individuals or species. It can be described as a variation (which may arise due to mutation or alteration in the genomic loci) that can be ...
s of 357 samples from captive and wild lions from Africa and India were examined. Results indicate four lineages of lion populations: one in Central and North Africa to Asia, one in
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
, one in Southern Africa, and one in Southern and
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historica ...
; the first wave of lion expansion probably occurred about 118,000 years ago from East Africa into
West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes ...
, and the second wave in the late Pleistocene or early
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
periods from Southern Africa towards East Africa. The Asiatic lion is genetically closer to North and West African lions than to the group comprising East and Southern African lions. The two groups probably diverged about 186,000–128,000 years ago. It is thought that the Asiatic lion remained connected to North and Central African lions until
gene flow In population genetics, gene flow (also known as gene migration or geneflow and allele flow) is the transfer of genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent a ...
was interrupted due to extinction of lions in Western Eurasia and the Middle East during the Holocene. Asiatic lions are less genetically diverse than African lions, which may be the result of a
founder effect In population genetics, the founder effect is the loss of genetic variation that occurs when a new population is established by a very small number of individuals from a larger population. It was first fully outlined by Ernst Mayr in 1942, us ...
in the recent history of the remnant population in the Gir Forest.


Characteristics

The Asiatic lion's fur ranges in colour from ruddy-tawny, heavily speckled with black, to sandy or buffish grey, sometimes with a silvery sheen in certain lighting. Males have only moderate mane growth at the top of the head, so that their ears are always visible. The mane is scanty on the cheeks and throat, where it is only long. About half of Asiatic lions' skulls from the Gir forest have divided infraorbital foramina, whereas African lions have only one foramen on either side. The
sagittal crest A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are exception ...
is more strongly developed, and the post-orbital area is shorter than in African lions. Skull length in adult males ranges from , and in females, from . It differs from the African lion by a larger tail tuft and less inflated
auditory bullae The tympanic part of the temporal bone is a curved plate of bone lying below the squamous part of the temporal bone, in front of the mastoid process, and surrounding the external part of the ear canal. It originates as a separate bone (tympanic b ...
. The most striking morphological character of the Asiatic lion is a longitudinal fold of skin running along its belly. Males have a shoulder height of up to , and females of . Two lions in Gir Forest measured from head to body with a long tail of and total lengths of . The Gir lion is similar in size to the Central African lion, and smaller than large African lions. An Adult male Asiatic lion weighs on average with the limit being ; a wild female weighs .


Manes

Colour and development of manes in male lions varies between regions, among populations and with age of lions. In general, the Asiatic lion differs from the African lion by a less developed mane. The manes of most lions in ancient Greece and Asia Minor were also less developed and did not extend to below the belly, sides or
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
s. Lions with such smaller manes were also known in the
Syrian region Syria (Hieroglyphic Luwian: 𔒂𔒠 ''Sura/i''; gr, Συρία) or Sham ( ar, ٱلشَّام, ash-Shām) is the name of a historical region located east of the Mediterranean Sea in Western Asia, broadly synonymous with the Levant. Other ...
,
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...
and
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a List of transcontinental countries, transcontinental country spanning the North Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via a land bridg ...
.


Exceptionally sized lions

The confirmed record total length of a male Asiatic lion is , including the tail. Emperor Jahangir allegedly speared a lion in the 1620s that measured and weighed . In 1841, English traveller
Austen Henry Layard Sir Austen Henry Layard (; 5 March 18175 July 1894) was an English Assyriologist, traveller, cuneiformist, art historian, draughtsman, collector, politician and diplomat. He was born to a mostly English family in Paris and largely raised in It ...
accompanied hunters in
Khuzestan Khuzestan Province (also spelled Xuzestan; fa, استان خوزستان ''Ostān-e Xūzestān'') is one of the 31 provinces of Iran. It is in the southwest of the country, bordering Iraq and the Persian Gulf. Its capital is Ahvaz and it cover ...
, Iran, and sighted a lion which "had done much damage in the plain of Ram Hormuz," before one of his companions killed it. He described it as being "unusually large and of very dark brown colour", with some parts of its body being almost black. In 1935, a British admiral claimed to have sighted a maneless lion near
Quetta Quetta (; ur, ; ; ps, کوټه‎) is the tenth most populous city in Pakistan with a population of over 1.1 million. It is situated in south-west of the country close to the International border with Afghanistan. It is the capital of th ...
in Pakistan. He wrote "It was a large lion, very stocky, light tawny in colour, and I may say that no one of us three had the slightest doubt of what we had seen until, on our arrival at Quetta, many officers expressed doubts as to its identity, or to the possibility of there being a lion in the district."


Distribution and habitat

In Saurashtra's Gir Forest, an area of was declared as a sanctuary for Asiatic lion conservation in 1965. This sanctuary and the surrounding areas are the only
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s supporting the Asiatic lion. After 1965, a national park was established covering an area of where human activity is not allowed. In the surrounding sanctuary only Maldharis have the right to take their livestock for grazing. Lions inhabit remnant forest habitats in the two hill systems of Gir and Girnar that comprise Gujarat's largest tracts of
tropical and subtropical dry broadleaf forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
, thorny forest and savanna, and provide valuable habitat for a diverse flora and fauna. Five protected areas currently exist to protect the Asiatic lion: Gir Sanctuary, Gir National Park, Pania Sanctuary, Mitiyala Sanctuary, and Girnar Sanctuary. The first three protected areas form the Gir Conservation Area, a large forest block that represents the core habitat of the lion population. The other two sanctuaries Mitiyala and Girnar protect satellite areas within dispersal distance of the Gir Conservation Area. An additional sanctuary is being established in the nearby
Barda Wildlife Sanctuary Barda Wildlife Sanctuary is located in Gujarat, India. It is situated approximately from Porbandar and west of Gir Forest National Park. Previous to its 1979 establishment as a wildlife sanctuary, Barda was a private reserve for Porbandar and ...
to serve as an alternative home for lions. The drier eastern part is vegetated with
acacia ''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus nam ...
thorn savanna and receives about annual rainfall; rainfall in the west is higher at about per year. The lion population recovered from the brink of
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the Endling, last individual of the species, although the Functional ext ...
to 411 individuals by 2010. In that year, approximately 105 lions lived outside the Gir forest, representing a quarter of the entire lion population. Dispersing sub-adults established new territories outside their natal prides, and as a result the satellite lion population has been increasing since 1995. By 2015, the total population had grown to an estimated 523 individuals, inhabiting an area of in the Saurashtra region. The Asiatic Lion Census conducted in 2017 revealed about 650 individuals. By 2020, at least six satellite populations had spread to eight districts in Gujarat and live in human-dominated areas outside the protected area network.


Former range

The Asiatic lion used to occur in
Arabia The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Pl ...
, Palestine, Mesopotamia and
Baluchistan Balochistan ( ; bal, بلۏچستان; also romanised as Baluchistan and Baluchestan) is a historical region in Western Asia, Western and South Asia, located in the Iranian plateau's far southeast and bordering the Indian Plate and the Arabian S ...
. In South Caucasia (present day
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''O ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
, and
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
), it was known since the
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
, and became extinct in the 10th century. Until the middle of the 19th century, it survived in regions adjoining Mesopotamia and
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
, and was still sighted in the upper reaches of the
Euphrates The Euphrates () is the longest and one of the most historically important rivers of Western Asia. Tigris–Euphrates river system, Together with the Tigris, it is one of the two defining rivers of Mesopotamia ( ''the land between the rivers'') ...
River in the early 1870s. By the late 19th century, the Asiatic lion had become extinct in
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the Ara ...
and
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
. The last known lion in
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
was killed on the lower Tigris in 1918. Historical records in Iran indicate that it ranged from the Khuzestan Plain to Fars Province at elevations below in
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate gras ...
vegetation and
pistachio The pistachio (, ''Pistacia vera''), a member of the cashew family, is a small tree originating from Central Asia and the Middle East. The tree produces seeds that are widely consumed as food. ''Pistacia vera'' is often confused with other spe ...
-
almond The almond (''Prunus amygdalus'', syn. ''Prunus dulcis'') is a species of tree native to Iran and surrounding countries, including the Levant. The almond is also the name of the edible and widely cultivated seed of this tree. Within the genu ...
woodlands. It was widespread in the country, but in the 1870s, it was sighted only on the western slopes of the
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgr ...
, and in the forest regions south of
Shiraz Shiraz (; fa, شیراز, Širâz ) is the fifth-most-populous city of Iran and the capital of Fars Province, which has been historically known as Pars () and Persis. As of the 2016 national census, the population of the city was 1,565,572 p ...
. It served as the national emblem and appeared on the country's flag. Some of the country's last lions were sighted in 1941 between Shiraz and Jahrom in Fars Province, and in 1942, a lion was spotted about northwest of
Dezful Dezful ( fa, دزفول, pronounced , Dezfuli dialect: Desfil, pronounced ) also Romanized as Dezfūl and Dezfool; also known as Dīzfūl and Ab I Diz is a city and capital of Dezful County, Khuzestan Province, Iran. At the 2011 census, its popu ...
. In 1944, the corpse of a lioness was found on the banks of the
Karun The Karun ( fa, کارون, ) is the Iranian river with the highest water flow, and its only navigable river. It is long. It rises in the Zard Kuh mountains of the Bakhtiari district in the Zagros Range, receiving many tributaries, such as t ...
River in Iran's
Khuzestan Province Khuzestan Province (also spelled Xuzestan; fa, استان خوزستان ''Ostān-e Xūzestān'') is one of the 31 provinces of Iran. It is in the southwest of the country, bordering Iraq and the Persian Gulf. Its capital is Ahvaz and it covers ...
. In India, the Asiatic lion occurred in
Sind Sindh (; ; ur, , ; historically romanized as Sind) is one of the four provinces of Pakistan. Located in the southeastern region of the country, Sindh is the third-largest province of Pakistan by land area and the second-largest province ...
,
Bahawalpur Bahawalpur () is a city in the Punjab province of Pakistan. With inhabitants as of 2017, it is Pakistan's 11th most populous city. Founded in 1748, Bahawalpur was the capital of the former princely state of Bahawalpur, ruled by the Abbasi fa ...
,
Punjab Punjab (; Punjabi Language, Punjabi: پنجاب ; ਪੰਜਾਬ ; ; also Romanization, romanised as ''Panjāb'' or ''Panj-Āb'') is a geopolitical, cultural, and historical region in South Asia, specifically in the northern part of the I ...
, Gujarat,
Rajasthan Rajasthan (; lit. 'Land of Kings') is a state in northern India. It covers or 10.4 per cent of India's total geographical area. It is the largest Indian state by area and the seventh largest by population. It is on India's northwestern ...
,
Haryana Haryana (; ) is an Indian state located in the northern part of the country. It was carved out of the former state of East Punjab on 1 Nov 1966 on a linguistic basis. It is ranked 21st in terms of area, with less than 1.4% () of India's land ...
,
Bihar Bihar (; ) is a state in eastern India. It is the 2nd largest state by population in 2019, 12th largest by area of , and 14th largest by GDP in 2021. Bihar borders Uttar Pradesh to its west, Nepal to the north, the northern part of West ...
and eastward as far as Palamau and Rewa, Madhya Pradesh in the early 19th century. It once ranged to Bangladesh in the east and up to Narmada River in the south. Because of the lion's restricted distribution in India,
Reginald Innes Pocock Reginald Innes Pocock Fellow of the Royal Society, F.R.S. (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock (historian), Nicholas Pocock and Edith Prichard. He ...
assumed that it arrived from Europe, southwestern Asia through Balochistan only recently, before humans started limiting its dispersal in the country. The advent and increasing availability of firearms led to its local extirpation over large areas. Heavy hunting by British colonial officers and
Indian rulers Indian or Indians may refer to: Peoples South Asia * Indian people, people of Indian nationality, or people who have an Indian ancestor ** Non-resident Indian, a citizen of India who has temporarily emigrated to another country * South Asia ...
caused a steady and marked decline of lion numbers in the country. Lions were exterminated in Palamau by 1814, in
Baroda Vadodara (), also known as Baroda, is the second largest city in the Indian state of Gujarat. It serves as the administrative headquarters of the Vadodara district and is situated on the banks of the Vishwamitri River, from the state capital ...
, Hariana and
Ahmedabad district {{Infobox settlement , name = Ahmedabad District , native_name = , native_name_lang = , settlement_type = District of Gujarat , image_skyline = D ...
in the 1830s, in
Kot Diji The ancient site at Kot Diji ( sd, ڪوٽ ڏیجي; ur, کوٹ ڈیجی) was the forerunner of the Indus Civilization. The occupation of this site is attested already at 3300 BCE. The remains consist of two parts; the citadel area on high ground ...
and Damoh in the 1840s. During the
Indian Rebellion of 1857 The Indian Rebellion of 1857 was a major uprising in India in 1857–58 against the rule of the British East India Company, which functioned as a sovereign power on behalf of the British Crown. The rebellion began on 10 May 1857 in the for ...
, a British officer shot 300 lions. The last lions of
Gwalior Gwalior() is a major city in the central Indian state of Madhya Pradesh; it lies in northern part of Madhya Pradesh and is one of the Counter-magnet cities. Located south of Delhi, the capital city of India, from Agra and from Bhopal, the s ...
and Rewah were shot in the 1860s. One lion was killed near
Allahabad Allahabad (), officially known as Prayagraj, also known as Ilahabad, is a metropolis in the Indian state of Uttar Pradesh.The other five cities were: Agra, Kanpur (Cawnpore), Lucknow, Meerut, and Varanasi (Benares). It is the administra ...
in 1866. The last lion of
Mount Abu Mount Abu () is a hill station in the Aravalli Range in Sirohi district of the state of Rajasthan in western India.The mountain forms a rocky plateau 22 km long by 9 km wide. The highest peak on the mountain is Guru Shikhar at abov ...
in
Rajasthan Rajasthan (; lit. 'Land of Kings') is a state in northern India. It covers or 10.4 per cent of India's total geographical area. It is the largest Indian state by area and the seventh largest by population. It is on India's northwestern ...
was spotted in 1872. By the late 1870s, lions were extinct in Rajasthan. By 1880, no lion survived in Guna, Deesa and
Palanpur Palanpur is a city and a municipality of Banaskantha district in the Indian state of Gujarat. Palanpur is the administrative headquarters of Banaskantha district. Palanpur is the ancestral home to an industry of Indian diamond merchants. Ety ...
districts, and only about a dozen lions were left in
Junagadh district Junagadh district is a district of the Indian state of Gujarat. Its administrative headquarters is the city of Junagadh. Geography The district is located on the Kathiawar peninsula in western Gujarat. It is surrounded by Rajkot District (N ...
. By the turn of the century, the Gir Forest held the only Asiatic lion population in India, which was protected by the
Nawab of Junagarh Nawab of Junagarh or Junagadh refers to the now defunct ex-lineage of rulers of the princely Junagarh State in British Raj, nowadays Junagadh district in the state of Gujarat in India. There are still several forts and palaces in India which wer ...
in his private hunting grounds.


Ecology and behaviour

Male Asiatic lions are solitary, or associate with up to three males, forming a loose
pride Pride is defined by Merriam-Webster as "reasonable self-esteem" or "confidence and satisfaction in oneself". A healthy amount of pride is good, however, pride sometimes is used interchangeably with "conceit" or "arrogance" (among other words) w ...
. Pairs of males rest, hunt and feed together, and display marking behaviour at the same sites. Females associate with up to 12 females, forming a stronger pride together with their cubs. They share large carcasses among each other, but seldom with males. Female and male lions usually associate only for a few days when
mating In biology, mating is the pairing of either opposite- sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reprod ...
, but rarely live and feed together. Results of a radio telemetry study indicate that annual home ranges of male lions vary from in dry and wet seasons. Home ranges of females are smaller, varying between . During hot and dry seasons, they favour densely vegetated and shady riverine habitats, where prey species also congregate. Coalitions of males defend home ranges containing one or more female prides. Together, they hold a territory for a longer time than single lions. Males in coalitions of three to four individuals exhibit a pronounced hierarchy with one male dominating the others. The lions in Gir National Park are active at twilight and by
night Night (also described as night time, unconventionally spelled as "nite") is the period of ambient darkness from sunset to sunrise during each 24-hour day, when the Sun is below the horizon. The exact time when night begins and ends depends ...
, showing a high temporal overlap with sambar (''Rusa unicolor''),
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species i ...
(''Sus scrofa'') and
nilgai The nilgai (''Boselaphus tragocamelus'') (, literally meaning "blue cow") is the largest Asian antelope and is ubiquitous across the northern Indian subcontinent. It is the sole member of the genus ''Boselaphus'' and was described by Peter Si ...
(''Boselaphus tragocamelus'').


Feeding ecology

In general, lions prefer large prey species within a weight range of , irrespective of their availability. Domestic cattle have historically been a major component of the Asiatic lions' diet in the Gir Forest. Inside Gir Forest National Park, lions predominantly kill chital (''Axis axis''), sambar deer, nilgai,
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult ...
(''Bos taurus''), domestic
water buffalo The water buffalo (''Bubalus bubalis''), also called the domestic water buffalo or Asian water buffalo, is a large bovid originating in the Indian subcontinent and Southeast Asia. Today, it is also found in Europe, Australia, North America, So ...
(''Bubalus bubalis''), and less frequently wild boar. They most commonly kill chital, which weighs only around . They prey on sambar deer when the latter descend from the hills during summer. Outside the protected area where wild prey species do not occur, lions prey on water buffalo and cattle, and rarely on
dromedary The dromedary (''Camelus dromedarius'' or ;), also known as the dromedary camel, Arabian camel, or one-humped camel, is a large even-toed ungulate, of the genus '' Camelus'', with one hump on its back. It is the tallest of the three species o ...
(''Camelus dromedarius''). They generally kill most prey less than away from water bodies, charge prey from close range and drag carcasses into dense cover. They regularly visit specific sites within the protected area to scavenge on dead livestock dumped by
Maldhari Maldharis are herdsmen community in Gujarat, India. Originally nomads, they came to be known as Maldharis after settling in Junagadh district (mainly Gir Forest). "Maldhari" is an occupational term which refers to people from a variety of castes ...
livestock herders. During dry, hot months, they also prey on
mugger crocodile The mugger crocodile (''Crocodylus palustris'') is a medium-sized broad-snouted crocodile, also known as mugger and marsh crocodile. It is native to freshwater habitats from southern Iran to the Indian subcontinent, where it inhabits marshe ...
s (''Crocodylus palustris'') on the banks of Kamleshwar Dam. In 1974, the Forest Department estimated the wild
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...
population at 9,650 individuals. In the following decades, the wild ungulate population has grown consistently to 31,490 in 1990 and 64,850 in 2010, including 52,490 chital, 4,440 wild boar, 4,000 sambar, 2,890 nilgai, 740
chinkara The chinkara (''Gazella bennettii''), also known as the Indian gazelle, is a gazelle species native to Iran, Afghanistan, Pakistan and India. Taxonomy The following six subspecies are considered valid: * Deccan chinkara (''G. b. bennettii ...
(''Gazella bennetti''), and 290 four-horned antelope (''Tetracerus quadricornis''). In contrast, populations of domestic buffalo and cattle declined following resettlement, largely due to direct removal of resident livestock from the Gir Conservation Area. The population of 24,250 domestic livestock in the 1970s declined to 12,500 by the mid-1980s, but increased to 23,440 animals in 2010. Following changes in both predator and prey communities, Asiatic lions shifted their predation patterns. Today, very few livestock kills occur within the sanctuary, and instead most occur in peripheral villages. Depredation records indicate that in and around the Gir Forest, lions killed on average 2,023 livestock annually between 2005 and 2009, and an additional 696 individuals in satellite areas. Dominant males consume about 47% more from kills than their coalition partners. Aggression between partners increases when coalitions are large, but kills are small.


Reproduction

Asiatic lions mate foremost between October and November. Mating lasts three to six days. During these days, they usually do not hunt, but only drink water.
Gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pr ...
lasts about 110 days. Litters comprise one to four cubs. The average interval between births is 24 months, unless cubs die due to
infanticide Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose is the prevention of resou ...
by adult males or because of diseases and injuries. Cubs become independent at the age of about two years. Subadult males leave their natal pride latest at the age of three years and become nomads until they establish their own territory. Dominant males mate more frequently than their coalition partners. During a study carried out between December 2012 and December 2016, three females were observed switching mating partners in favour of the dominant male. Monitoring of more than 70 mating events showed that females mated with males of several rivaling prides that shared their home ranges, and that these males were tolerant toward the same cubs. Only new males that entered the female territories killed unfamiliar cubs. Young females mated foremost with males within their home ranges. Older females selected males at the periphery of their home ranges.


Threats

The Asiatic lion currently exists as a single subpopulation, and is thus vulnerable to extinction from unpredictable events, such as an
epidemic An epidemic (from Greek ἐπί ''epi'' "upon or above" and δῆμος ''demos'' "people") is the rapid spread of disease to a large number of patients among a given population within an area in a short period of time. Epidemics of infectious ...
or large forest fire. There are indications of poaching incidents in recent years, as well as reports that organized poacher gangs have switched attention from local Bengal tigers to the Gujarat lions. There have also been a number of drowning incidents, after lions fell into wells. Prior to the resettlement of Maldharis, the Gir forest was heavily degraded and used by livestock, which competed with and restricted the population sizes of native ungulates. Various studies reveal tremendous habitat recovery and increases in wild ungulate populations following the resettlement of Maldharis since the 1970s. Nearly 25 lions in the vicinity of Gir Forest were found dead in October 2018. Four of them had died because of canine distemper virus, the same virus that had also killed several lions in the
Serengeti The Serengeti ( ) ecosystem is a geographical region in Africa, spanning northern Tanzania. The protected area within the region includes approximately of land, including the Serengeti National Park and several game reserves. The Serenget ...
.


Conflicts with humans

Since the mid 1990s, the Asiatic lion population has increased to an extent that by 2015, about a third resided outside the protected area. Hence, conflict between local residents and wildlife also increased. Local people protect their crops from nilgai, wild boar, and other
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
s by using electrical fences that are powered with high voltage. Some consider the presence of predators a benefit, as they keep the herbivore population in check. But some also fear the lions, and killed several in retaliation for attacks on livestock. In July 2012, a lion dragged a man from the veranda of his house and killed him about from Gir Forest National Park. This was the second attack by a lion in this area, six months after a 25-year-old man was attacked and killed in Dhodadar.


Conservation

''Panthera leo persica'' was included on CITES Appendix I, and is fully protected in India, where it is considered
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
.


Reintroduction


India

In the 1950s, biologists advised the Indian government to re-establish at least one wild population in the Asiatic lion's former range to ensure the population's reproductive health and to prevent it from being affected by an outbreak of an
epidemic An epidemic (from Greek ἐπί ''epi'' "upon or above" and δῆμος ''demos'' "people") is the rapid spread of disease to a large number of patients among a given population within an area in a short period of time. Epidemics of infectious ...
. In 1956, the Indian Board for Wildlife accepted a proposal by the Government of
Uttar Pradesh Uttar Pradesh (; , 'Northern Province') is a state in northern India. With over 200 million inhabitants, it is the most populated state in India as well as the most populous country subdivision in the world. It was established in 195 ...
to establish a new sanctuary for the envisaged reintroduction, Chandra Prabha Wildlife Sanctuary, covering in eastern Uttar Pradesh, where climate, terrain and vegetation is similar to the conditions in the Gir Forest. In 1957, one male and two female wild-caught Asiatic lions were set free in the sanctuary. This population comprised 11 animals in 1965, which all disappeared thereafter. The
Asiatic Lion Reintroduction Project The Asiatic Lion Reintroduction Project is an initiative of the Indian Government to provide safeguards to the Asiatic lion (''Panthera leo leo'') from extinction in the wild by means of reintroduction. The last wild population of the Asiatic lion ...
to find an alternative habitat for reintroducing Asiatic lions was pursued in the early 1990s. Biologists from the
Wildlife Institute of India The Wildlife Institute of India (WII) is an autonomous natural resource service institution established in 1982 under the Ministry of Environment Forest and Climate change, Government of India. WII carries out wildlife research in areas of stu ...
assessed several potential translocation sites for their suitability regarding existing prey population and habitat conditions. The Palpur-
Kuno Wildlife Sanctuary Kuno National Park is national park and Wildlife Sanctuary in Madhya Pradesh, India. It derives its name from Kuno River. It was established in 1981 as a wildlife sanctuary with an initial area of in the Sheopur and Morena districts. In 2018 ...
in northern Madhya Pradesh was ranked as the most promising location, followed by Sita Mata Wildlife Sanctuary and
Darrah National Park Mukundara Hills National Park is a national park in Rajasthan, India with an area of . It was established in 2004 and consists of three wildlife sanctuaries: Darrah Wildlife Sanctuary, National Chambal Sanctuary, and Jawahar Sagar Wildlife San ...
. Until 2000, 1,100 families from 16 villages had been resettled from the Palpur-Kuno Wildlife Sanctuary, and another 500 families from eight villages were expected to be resettled. With this resettlement scheme the protected area was expanded by . Gujarat state officials resisted the relocation, since it would make the Gir Sanctuary lose its status as the world's only home of the Asiatic lion. Gujarat raised a number of objections to the proposal, and thus the matter went before the
Indian Supreme Court The Supreme Court of India (IAST: ) is the supreme judicial authority of India and is the highest court of the Republic of India under the constitution. It is the most senior constitutional court, has the final decision in all legal matters e ...
. In April 2013, the Indian Supreme Court ordered the Gujarat state to send some of their Gir lions to Madhya Pradesh to establish a second population there. The court had given wildlife authorities six months to complete the transfer. The number of lions and which ones to be transported will be decided at a later date. As of now, the plan to shift lions to Kuno is in jeopardy, with Madhya Pradesh having apparently given up on acquiring lions from Gujarat.


Iran

In 1977, Iran attempted to restore its lion population by transporting Gir lions to Arzhan National Park, but the project met resistance from the local population, and thus it was not implemented. However, this did not stop Iran from seeking to bring back the lion. In February 2019,
Tehran Zoological Garden The Tehran Zoological Garden ( fa, ), also known as "Eram Zoo" (), is located on the Tehran–Karaj Freeway in the Eram Amusement Park, Tehran province, Iran. Inaugurated in 1992, the zoo covers an area of and houses over 290 species. Histo ...
obtained a male Asiatic lion from
Bristol Zoo Bristol Zoo was a zoo in the city of Bristol in South West England. The zoo's stated mission was to "maintain and defend” biodiversity through breeding endangered species, conserving threatened species and habitats and promoting a wider unders ...
in the United Kingdom, followed in June by a female from
Dublin Zoo Dublin Zoo ( ga, Zú Bhaile Átha Cliath), in Phoenix Park, Dublin, is a zoo in Ireland, and one of Dublin's most popular attractions. Established and designed in 1830 by Decimus Burton, it opened the following year. Today it focuses on conserv ...
. There are hopes for them to successfully reproduce.


In captivity

Until the late 1990s, captive Asiatic lions in Indian zoos were haphazardly interbred with African lions confiscated from circuses, leading to
genetic pollution Genetic pollution is a controversial term for uncontrolled gene flow into wild populations. It is defined as "the dispersal of contaminated altered genes from genetically engineered organisms to natural organisms, esp. by cross-pollination", but ...
in the captive Asiatic lion stock. Once discovered, this led to the complete shutdown of the European and American endangered species breeding programs for Asiatic lions, as its founder animals were captive-bred Asiatic lions originally imported from India and were ascertained to be intraspecific hybrids of African and Asian lions. In North American zoos, several Indian-African lion crosses were inadvertently bred, and researchers noted that "the fecundity, reproductive success, and spermatozoal development improved dramatically."
DNA fingerprinting DNA profiling (also called DNA fingerprinting) is the process of determining an individual's DNA characteristics. DNA analysis intended to identify a species, rather than an individual, is called DNA barcoding. DNA profiling is a forensic t ...
studies of Asiatic lions have helped in identifying individuals with high genetic variability, which can be used for conservation breeding programs. In 2006, the Central Zoo Authority of India stopped breeding Indian-African cross lions stating that "hybrid lions have no conservation value and it is not worth to spend resources on them". Now only pure native Asiatic lions are bred in India. In 1972 the Sakkarbaug Zoo sold a pair of young pure-stock lions to the Fauna Preservation Society; which decided they would be accommodated at the Jersey Wildlife Trust where it was hoped to begin a captive breeding programme. The Asiatic lion International Studbook was initiated in 1977, followed in 1983 by the
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
n
Species Survival Plan The American Species Survival Plan or SSP program was developed in 1981 by the (American) Association of Zoos and Aquariums to help ensure the survival of selected species in zoos and aquariums, most of which are threatened or endangered in the ...
(SSP). The North American population of captive Asiatic lions was composed of descendants of five founder lions, three of which were pure Asian and two were African or African-Asian hybrids. The lions kept in the framework of the SSP consisted of animals with high
inbreeding Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genetic disorders a ...
coefficients. In the early 1990s, three European zoos imported pure Asiatic lions from India:
London Zoo London Zoo, also known as ZSL London Zoo or London Zoological Gardens is the world's oldest scientific zoo. It was opened in London on 27 April 1828, and was originally intended to be used as a collection for scientific study. In 1831 or 1832, ...
obtained two pairs; the
Zürich Zoologischer Garten The Zoo Zürich is a zoo located in Zürich, Switzerland and together with the Basel Zoo is considered one of the best zoos in Europe. Opened in 1929, it is the third oldest zoo in Switzerland (after Basel and Arth-Goldau) and it accumulated ...
one pair; and the Korkeasaari Zoo in
Helsinki Helsinki ( or ; ; sv, Helsingfors, ) is the Capital city, capital, primate city, primate, and List of cities and towns in Finland, most populous city of Finland. Located on the shore of the Gulf of Finland, it is the seat of the region of U ...
one male and two females. In 1994, the
European Endangered Species Programme The EAZA Ex-situ Programme (EEP) is a population management and conservation programme by European Association of Zoos and Aquaria (EAZA) for wild animals living in European zoos. The programme was formerly known as the European Endangered Species ...
(EEP) for Asiatic lions was initiated. The European Association of Zoos and Aquaria (EAZA) published the first European Studbook in 1999. By 2005, there were 80 Asiatic lions kept in the EEP — the only captive population outside of India. As of 2009, more than 100 Asiatic lions were kept within the EEP. The SSP had not resumed; pure-bred Asiatic lions are needed to form a new founder population for breeding in American zoos.


In culture


South and East Asia

Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several pa ...
cave paintings of lions were found in Bhimbetka rock shelters in central India, which are at least 30,000 years old. The
Sanskrit Sanskrit (; attributively , ; nominally , , ) is a classical language belonging to the Indo-Aryan languages, Indo-Aryan branch of the Indo-European languages. It arose in South Asia after its predecessor languages had Trans-cultural diffusion ...
word for 'lion' is , which is also a name of
Shiva Shiva (; sa, शिव, lit=The Auspicious One, Śiva ), also known as Mahadeva (; Help:IPA/Sanskrit, ɐɦaːd̪eːʋɐ, or Hara, is one of the Hindu deities, principal deities of Hinduism. He is the Supreme Being in Shaivism, one o ...
and signifies the '' Leo'' of the
Zodiac The zodiac is a belt-shaped region of the sky that extends approximately 8° north or south (as measured in celestial latitude) of the ecliptic, the apparent path of the Sun across the celestial sphere over the course of the year. The pa ...
. The Sanskrit name of
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
is Sinhala meaning 'Abode of Lions'.
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, bor ...
derives its name from the Malay words 'lion' and 'city', which in turn is from the Sanskrit and , latter also meaning 'fortified town'. In
Hindu mythology Hindu mythology is the body of myths and literature attributed to, and espoused by, the adherents of the Hindu religion, found in Hindu texts such as the Vedic literature, epics like ''Mahabharata'' and ''Ramayana'', the Puranas, and ...
, the half man half lion
avatar Avatar (, ; ), is a concept within Hinduism that in Sanskrit literally means "descent". It signifies the material appearance or incarnation of a powerful deity, goddess or spirit on Earth. The relative verb to "alight, to make one's appear ...
Narasimha Narasimha ( sa, नरसिंह, lit=man-lion, ), sometimes rendered Narasingha, is the fourth avatar of the Hindu god Vishnu. He is regarded to have incarnated in the form of a part-lion, part-man being to slay Hiranyakashipu, to end rel ...
is the fourth
incarnation Incarnation literally means ''embodied in flesh'' or ''taking on flesh''. It refers to the conception and the embodiment of a deity or spirit in some earthly form or the appearance of a god as a human. If capitalized, it is the union of divinit ...
of
Vishnu Vishnu ( ; , ), also known as Narayana and Hari, is one of the principal deities of Hinduism. He is the supreme being within Vaishnavism, one of the major traditions within contemporary Hinduism. Vishnu is known as "The Preserver" withi ...
. Simhamukha is a lion-faced protector and
dakini A ḍākinī ( sa, डाकिनी; ; mn, хандарма; ; alternatively 荼枳尼, ; 荼吉尼, ; or 吒枳尼, ; Japanese: 荼枳尼 / 吒枳尼 / 荼吉尼, ''dakini'') is a type of female spirit, goddess, or demon in Hinduism and Bud ...
in
Tibetan Buddhism Tibetan Buddhism (also referred to as Indo-Tibetan Buddhism, Lamaism, Lamaistic Buddhism, Himalayan Buddhism, and Northern Buddhism) is the form of Buddhism practiced in Tibet and Bhutan, where it is the dominant religion. It is also in majo ...
. In the 18th book of the
Mahabharata The ''Mahābhārata'' ( ; sa, महाभारतम्, ', ) is one of the two major Sanskrit epics of ancient India in Hinduism, the other being the '' Rāmāyaṇa''. It narrates the struggle between two groups of cousins in the K ...
, Bharata deprives lions of their prowess. The lion plays a prominent role in '' The Fables of Pilpay'' that were translated into Persian, Greek and
Hebrew language Hebrew (; ; ) is a Northwest Semitic language of the Afroasiatic language family. Historically, it is one of the spoken languages of the Israelites and their longest-surviving descendants, the Jews and Samaritans. It was largely preserve ...
s between the 8th and 12th centuries. The lion is the symbol of
Mahavira Mahavira (Sanskrit: महावीर) also known as Vardhaman, was the 24th ''tirthankara'' (supreme preacher) of Jainism. He was the spiritual successor of the 23rd ''tirthankara'' Parshvanatha. Mahavira was born in the early part of the 6 ...
, the 24th and last
Tirthankara In Jainism, a ''Tirthankara'' (Sanskrit: '; English: literally a ' ford-maker') is a saviour and spiritual teacher of the '' dharma'' (righteous path). The word ''tirthankara'' signifies the founder of a '' tirtha'', which is a fordable pass ...
in
Jainism Jainism ( ), also known as Jain Dharma, is an Indian religion. Jainism traces its spiritual ideas and history through the succession of twenty-four tirthankaras (supreme preachers of ''Dharma''), with the first in the current time cycle bein ...
. *The lion is the third animal of the
Burmese zodiac The Burmese zodiac ( my, ဇာတာ ရာသီခွင် ) is the traditional Burmese system of astronomy and astrology. While it is still an important component of the Burmese calendar, today, the zodiac is closely identified with Burmese ...
and the sixth animal of the Sinhalese zodiac. *The earliest known Chinese stone sculptures of lions date to the
Han dynasty The Han dynasty (, ; ) was an Dynasties in Chinese history, imperial dynasty of China (202 BC – 9 AD, 25–220 AD), established by Emperor Gaozu of Han, Liu Bang (Emperor Gao) and ruled by the House of Liu. The dynasty was preceded by th ...
at the turn of the first millennium. *The
lion dance F Lion dance () is a form of traditional dance in Chinese culture and other Asian countries in which performers mimic a lion's movements in a lion costume to bring good luck and fortune. The lion dance is usually performed during the Chinese New Y ...
is a traditional dance in
Chinese culture Chinese culture () is one of the world's oldest cultures, originating thousands of years ago. The culture prevails across a large geographical region in East Asia and is extremely diverse and varying, with customs and traditions varying grea ...
that is strongly associated with Buddhism and known since at least the Han dynasty. *
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand ...
has a native martial art called '' Bokator'' ( km, ល្បុក្កតោ, pounding a lion).


West Asia and Europe

Lions are depicted on vases dating to about 2600
before present Before Present (BP) years, or "years before present", is a time scale used mainly in archaeology, geology and other scientific disciplines to specify when events occurred relative to the origin of practical radiocarbon dating in the 1950s. Beca ...
that were excavated near Lake Urmia in Iran. The lion was an important symbol in
Ancient Iraq Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the ...
and is depicted in a stone relief at
Nineveh Nineveh (; akk, ; Biblical Hebrew: '; ar, نَيْنَوَىٰ '; syr, ܢܝܼܢܘܹܐ, Nīnwē) was an ancient Assyrian city of Upper Mesopotamia, located in the modern-day city of Mosul in northern Iraq. It is located on the eastern ba ...
in the Mesopotamian Plain. * The lion makes repeated appearances in the
Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity Christianity is an Abrahamic monotheistic religion based on the life and teachings of Jesus ...
, most notably as having fought Samson in the
Book of Judges The Book of Judges (, ') is the seventh book of the Hebrew Bible and the Christian Old Testament. In the narrative of the Hebrew Bible, it covers the time between the conquest described in the Book of Joshua and the establishment of a kingdo ...
. * Having occurred in the
Arab world The Arab world ( ar, اَلْعَالَمُ الْعَرَبِيُّ '), formally the Arab homeland ( '), also known as the Arab nation ( '), the Arabsphere, or the Arab states, refers to a vast group of countries, mainly located in Western A ...
, particularly the Arabian Peninsula, the Asiatic lion has significance in
Arab The Arabs (singular: Arab; singular ar, عَرَبِيٌّ, DIN 31635: , , plural ar, عَرَب, DIN 31635: , Arabic pronunciation: ), also known as the Arab people, are an ethnic group mainly inhabiting the Arab world in Western Asia, ...
and
Islamic culture Islamic culture and Muslim culture refer to cultural practices which are common to historically Islamic people. The early forms of Muslim culture, from the Rashidun Caliphate to the early Umayyad period and the early Abbasid period, were predom ...
. For example, Surat al-Muddaththir of the
Quran The Quran (, ; Standard Arabic: , Quranic Arabic: , , 'the recitation'), also romanized Qur'an or Koran, is the central religious text of Islam, believed by Muslims to be a revelation from God. It is organized in 114 chapters (pl.: , ...
criticizes people who were averse to the
Islamic Prophet Prophets in Islam ( ar, الأنبياء في الإسلام, translit=al-ʾAnbiyāʾ fī al-ʾIslām) are individuals in Islam who are believed to spread God's message on Earth and to serve as models of ideal human behaviour. Some prophets ar ...
Muhammad Muhammad ( ar, مُحَمَّد;  570 – 8 June 632 CE) was an Arab religious, social, and political leader and the founder of Islam. According to Islamic doctrine, he was a prophet divinely inspired to preach and confirm the mon ...
's teachings, such as that the rich have an obligation to donate wealth to the poor, comparing their attitude to itself, with the response of prey to a ''qaswarah'' ( ar, قَسْوَرَة, meaning "lion", "beast of prey", or "hunter"). Other Arabic words for 'lion' include ''asad'' ( ar, أَسَد) and ''sabaʿ'' ( ar, سَبَع), and they can be used as names of places, or titles of people. An Arabic toponym for the
Levant The Levant () is an approximate historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology and other cultural contexts, it is ...
ine City of
Beersheba Beersheba or Beer Sheva, officially Be'er-Sheva ( he, בְּאֵר שֶׁבַע, ''Bəʾēr Ševaʿ'', ; ar, بئر السبع, Biʾr as-Sabʿ, Well of the Oath or Well of the Seven), is the largest city in the Negev desert of southern Israel. ...
( ar, بِئر ٱلسَّبَع) can mean "Spring of the Lion." Hamzah ibn Abdul-Muttalib who were loyal kinsmen of Muhammad, were given titles like ''Asad
Allah Allah (; ar, الله, translit=Allāh, ) is the common Arabic word for God. In the English language, the word generally refers to God in Islam. The word is thought to be derived by contraction from '' al- ilāh'', which means "the god", a ...
'' ( ar, أَسَد ٱلله, lit=Lion of God). * The lion of Babylon is a statue at the
Ishtar Gate The Ishtar Gate was the eighth gate to the inner city of Babylon (in the area of present-day Hillah, Babil Governorate, Iraq). It was constructed circa 575 BCE by order of King Nebuchadnezzar II on the north side of the city. It was pa ...
in
Babylon ''Bābili(m)'' * sux, 𒆍𒀭𒊏𒆠 * arc, 𐡁𐡁𐡋 ''Bāḇel'' * syc, ܒܒܠ ''Bāḇel'' * grc-gre, Βαβυλών ''Babylṓn'' * he, בָּבֶל ''Bāvel'' * peo, 𐎲𐎠𐎲𐎡𐎽𐎢 ''Bābiru'' * elx, 𒀸𒁀𒉿𒇷 ''Babi ...
The lion has an important association with the figure
Gilgamesh sux, , label=none , image = Hero lion Dur-Sharrukin Louvre AO19862.jpg , alt = , caption = Possible representation of Gilgamesh as Master of Animals, grasping a lion in his left arm and snake in his right hand, in an Assy ...
, as demonstrated in his epic. The Iraqi national football team is nicknamed "Lions of Mesopotamia." * The symbol of the lion is closely tied to the
Persian people The Persians are an Iranian ethnic group who comprise over half of the population of Iran. They share a common cultural system and are native speakers of the Persian language as well as of the languages that are closely related to Persian. ...
.
Achaemenid The Achaemenid Empire or Achaemenian Empire (; peo, wikt:𐎧𐏁𐏂𐎶, 𐎧𐏁𐏂, , ), also called the First Persian Empire, was an History of Iran#Classical antiquity, ancient Iranian empire founded by Cyrus the Great in 550 BC. Bas ...
kings were known to carry the symbol of the lion on their thrones and garments. The name 'Shir' (also pronounced 'Sher') ( fa, ) is a part of the names of many places in Iran and Central Asia, like those of city of
Shiraz Shiraz (; fa, شیراز, Širâz ) is the fifth-most-populous city of Iran and the capital of Fars Province, which has been historically known as Pars () and Persis. As of the 2016 national census, the population of the city was 1,565,572 p ...
and the Sherabad River, and had been adopted into other languages, like
Hindi Hindi (Devanāgarī: or , ), or more precisely Modern Standard Hindi (Devanagari: ), is an Indo-Aryan language spoken chiefly in the Hindi Belt region encompassing parts of northern, central, eastern, and western India. Hindi has been ...
. The ''Shir-va-Khorshid'' ( fa, , "
Lion and Sun The Lion and Sun ( fa, شیر و خورشید, Šir-o xoršid, ; Classical Persian: ) is one of the main emblems of Iran (Persia), and was an element in Iran's national flag until the 1979 Iranian Revolution and is still commonly used by oppo ...
") is one of the most prominent symbols of Iran, dating back to the
Safavid dynasty The Safavid dynasty (; fa, دودمان صفوی, Dudmâne Safavi, ) was one of Iran's most significant ruling dynasties reigning from 1501 to 1736. Their rule is often considered the beginning of modern Iranian history, as well as one of th ...
, and was used on the flag of Iran until 1979. * The lion was an objective of
hunting Hunting is the human activity, human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products (fur/hide (skin), hide, ...
in the Caucasus, by both locals and foreigners. The locals were called ' Shirvanshakhs'. * The Nemean lion of pre-literate Greek myth is associated with the Labours of Herakles. * A
Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...
statue of a lion from either
Southern Italy Southern Italy ( it, Sud Italia or ) also known as ''Meridione'' or ''Mezzogiorno'' (), is a macroregion of the Italian Republic consisting of its southern half. The term ''Mezzogiorno'' today refers to regions that are associated with the pe ...
or southern
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
from around 1000–1200 years Before Christ, the "Mari-Cha Lion", was exhibited at the Louvre Abu Dhabi.


See also

* Lion populations: Cape lion Lions in Europe *
Wildlife of Iran The wildlife of Iran include the fauna and flora of Iran. One of the most famous wildlife of Iran is the critically endangered Asiatic cheetah (''Acinonyx jubatus venaticus''), which today survives only in Iran. History The wildlife of Iran first ...
*
Wildlife of India India is home to a large variety of wildlife. It is a biodiversity hotspot with its various ecosystems ranging from the Himalayas in the north to the evergreen rain forests in the south, the sands of the west to the marshy mangroves of the ea ...
* ''In situ'' conservation * ''Ex situ'' conservation * '' Panthera leo fossilis'' * '' Panthera spelaea'' * '' Damnatio ad bestias''


References


Further reading

* * * * *


External links


IUCN/SSC Cat Specialist Group: Asiatic lionThe Telegraph, August 2018: Pride of India
*
Asiatic Lion Protection Society (ALPS), Gujarat, India





Asiatic lions in online video
(3 videos)
Asiatic Lions Images
* by Rajesh Badal (2011)
DB Video Special Report on Asiatic lion in Gujarati: What Is the connection Between Gir lions and Africans lions

Skin of a Persian lioness, belonging to an endangered subspecies of lions
brought to
Dublin Dublin (; , or ) is the capital and largest city of Ireland. On a bay at the mouth of the River Liffey, it is in the province of Leinster, bordered on the south by the Dublin Mountains, a part of the Wicklow Mountains range. At the 2016 ...
by King
Edward VII Edward VII (Albert Edward; 9 November 1841 – 6 May 1910) was King of the United Kingdom of Great Britain and Ireland and Emperor of India, from 22 January 1901 until Death and state funeral of Edward VII, his death in 1910. The second chil ...
in 1902 (during the reign of Shah Mozaffar ad-Din in
Persia Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
, and kept in the
Natural History Museum (Ireland) Natural History Museum primarily refers to: * Natural history museum (in title case), a scientific institution with natural history collections Natural History Museum may also refer to: * Natural History Museum, Abu Dhabi * Natural History Museum, ...
).
Lion of Basrah

A lion in Iraq

Stuffed animals
including Pakistan's last wild lion at Bahawalpur Zoo

of Muhammad's descendant
Musa al-Kadhim Musa ibn Ja'far al-Kazim ( ar, مُوسَىٰ ٱبْن جَعْفَر ٱلْكَاظِم, Mūsā ibn Jaʿfar al-Kāẓim), also known as Abū al-Ḥasan, Abū ʿAbd Allāh or Abū Ibrāhīm, was the seventh Imam in Twelver Shia Islam, after hi ...
encountering a lion outside
Medina Medina,, ', "the radiant city"; or , ', (), "the city" officially Al Madinah Al Munawwarah (, , Turkish: Medine-i Münevvere) and also commonly simplified as Madīnah or Madinah (, ), is the Holiest sites in Islam, second-holiest city in Islam, ...
in the
mountainous A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited summit area, and is usually higher th ...
region of the
Hejaz The Hejaz (, also ; ar, ٱلْحِجَاز, al-Ḥijāz, lit=the Barrier, ) is a region in the west of Saudi Arabia. It includes the cities of Mecca, Medina, Jeddah, Tabuk, Yanbu, Taif, and Baljurashi. It is also known as the "Western Prov ...

Description of the Arabian lion and art

4 انواع الأسود في العالم الأسد العربي الجزء
(in
Arabic Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walter ...
)
الاسد العربي المنقرض عند العرب lion Arabian Extinct
(in Arabic)
Asiatic lioness on a tree
{{Taxonbar, from=Q182347 Articles containing video clips Felids of Asia Felids of India Fauna of Gujarat Panthera leo leo Mammals described in 1826 Symbols of Gujarat