Arcticodactylus

''Arcticodactylus'' is a genus of basal pterosaur living during the Late Triassic in the area of present Greenland. Its only species was previously attributed to ''Eudimorphodon'', and its closest relatives may have been ''Eudimorphodon'' or ''Austriadraco''.

History of discovery

In 1989, William Amaral on the McKnight Bjerg in the east of Greenland discovered a rich fossil site. It was excavated in 1991 and 1992. Part of the material was a small skeleton of a pterosaur. In 2001, Farish Jenkins, Neil Shubin, Stephen Gatesy and Kevin Padian named and described it as a new species of ''Eudimorphodon'': ''Eudimorphodon cromptonellus''. The specific name (zoology), specific name honors Professor Alfred Walter Crompton. The suffix ''~ellus'', in Latin indicating a diminutive, alluded to the small size of the specimen.

The holotype, MGUH VP 3393, was found in the Carlsberg Fjord Beds of the Ørsted Dal Member of the Fleming Fjord Formation dating from the Norian – Rhaetian. It consists of a partial skeleton with skull. It is largely disarticulated.

The reference to ''Eudimorphodon'' had been essentially based on the similarity in tooth form, especially the distinctive multi-cuspid build with three, four or five points on the crown. In 2003, Alexander Kellner pointed out that other basal pterosaurs also possess such teeth. In 2014, Fabio Marco Dalla Vecchia noted that ''E. cromptonellus'' shared not a single trait with ''Eudimorphodon ranzii'' not present in other pterosaurs but lacked the distinguishing fang-like teeth, pterygoid teeth and striated tooth enamel. In 2015, Kellner named a separate genus ''Arcticodactylus''. The generic name is derived from the Arctic, and Greek δάκτυλος, ''daktylos'', "finger", a usual suffix in pterosaur names since ''Pterodactylus''. The ''Life Science Identifiers, Life Science Identifier'' is 72AE012A-018A-4B4B-950F-3CCB4C1D2471. The type species is ''Eudimorphodon cromptonellus'', the ''combinatio nova'' is ''Arcticodactylus cromptonellus''.

Description

The holotype individual of ''Arcticodactylus'' is the smallest pterosaur known, with an estimated wingspan of just . It was in 2001, on the basis of histological research of its bone structure, considered not to have been full-grown yet, though not newly born.

In 2015, Kellner established some distinguishing traits, correcting and adding to the 2001 diagnosis. The jaws have eleven or twelve multi-cusped teeth per side. The articulation surface of the fourth metacarpal with the fourth finger shows two true condyles. The thighbone is only a little shorter than the shinbone, with 96% of its length. The scapula is much longer, 93%, than the coracoid. The humerus is only slightly shorter than the thighbone, with 92% of its length, or the ulna with 91% of ulnar length. The thighbone is somewhat longer than the first phalanx of the wing finger that has 91% of femoral length. The third metatarsal of the foot is elongated with 56% of shinbone length. These proportions imply that ''Arcticodactylus'' had relatively short wings and large feet.

''Arcticodactylus'' can furthermore be distinguished from ''Eudimorphodon'' in the lack of long fang-like teeth in the middle of the tooth row and from ''Eudimorphodon ranzii'', ''Carniadactylus'' and ''Bergamodactylus'' by a triangular instead of rectangular deltopectoral crest on the humerus. ''Articodactylus'' has fewer teeth than any other known Triassic pterosaur.

Jenkins e.a. claimed that the unique articulation in ''Arcticodactylus'' between the main wing metacarpal and the wing finger, with two rounded condyles, was a transitional shape between the ancestral form that featured a single rounded articulation surface on the metacarpal allowing a considerable amount of lateral movement, and the condition in later pterosaurs that showed a gentle depression or trochlea. The two condyles, the upper one the largest, would have forced the finger into the most optimal plane of movement during the upstroke of the wing.

Classification

In 2001, ''E. cromptonellus'' was placed in the Eudimorphodontidae. Kellner in 2015 indicated a basal position in the Pterosauria, the short coracoid suggesting a close affinity to ''Austriadraco'' within an Austriadraconidae. According to Kellner, the original describers had incorrectly identified a coracoid as a quadrate bone. The following phylogenetic analysis follows the topology of Upchurch ''et al.'' (2015).

In 2020 however, a study upheld by Matthew G. Baron about early pterosaur interrelationships found ''Arcticodactylus'' to group with ''Carniadactylus'', ''Raeticodactylus'', and the Austriadraconidae, which in turn were within a clade he called Caviramidae.Matthew G. Baron (2020). "Testing pterosaur ingroup relationships through broader sampling of avemetatarsalian taxa and characters and a range of phylogenetic analysis techniques". PeerJ. 8: e9604. doi:10.7717/peerj.9604. PMC 7512134. .

See also

* List of pterosaur genera

References

{{Taxonbar, from=Q20651334

Late Triassic pterosaurs of Europe
Pterosaurs
Fossil taxa described in 2015