Anteosauridae is an extinct

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

of large

carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...

dinocephalia Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephal ...

therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...

s that are known from the Middle Permian of Asia, Africa, and South America.These animals were by far the largest predators of the

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleo ...

period, with skulls reaching 80 cm in length in adult individuals, far larger than the biggest

gorgonopsia Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, a ...

Description

Anteosaurids are characterized by very large pointed

incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...

s and canines, bulbous spatulate (spoon shaped) postcanines, a very strongly upturned margin of the

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...

, so the front of mouth curves strongly upwards, and a long, very robust lower jaw (Boonstra 1963). Anteosaurids distinguished from '' Brithopus'' and other non-anteosaurid anteosaurians by the presence of a large thickened region or "boss" on the side of the angle of the lower jaw; this was probably used in intraspecific behaviour. In ''

Doliosauriscus ''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneu ...

'' and ''

Anteosaurus ''Anteosaurus'' (meaning "Antaeus reptile") is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian (= Middle Permian) during the Capitanian stage, about 265 to 260 million years ago in what is now ...

'', not only was this boss very prominent, but the bones were very thick and rugose. The same situation is found in the herbivorous

Tapinocephalidae Tapinocephalidae was an advanced family of tapinocephalians. It is defined as the clade containing '' Ulemosaurus'', ''Tapinocaninus'', and the Tapinocephalinae. They are known from both Russia and South Africa. In all probability, the Tapinocep ...

, and it has been proposed that these animals engaged in head-butting behaviour. (Hopson & Barghusen 1986)

Classification

The subfamily

Anteosaurinae Anteosaurinae is an extinct subfamily of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Syodontinae. Description These are very specialized, very large anteosaurs. The postcanine teeth are f ...

has also been used in the past to include these therapsids. The Russian genera were previously included by Efremov in the family

Brithopodidae ''Brithopus'' is an extinct genus of dinocephalian therapsids. It contains a single species, ''Brithopus priscus'', known from fragmentary remains found in the Copper Sandstones near Isheevo, Russia. Description ''Brithopus'' was fairly large, ...

. Anteosauridae is part of a larger group of dinocephalians called

Anteosauria Anteosaurs are a group of large, primitive carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls ...

. Several recent

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...

studies of anteosaurians find support for two subfamilies within Anteosauridae: Anteosaurinae and

Syodontinae Syodontinae is a group of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Anteosaurinae. They are known from the Middle Permian Period of what is now Russia and South Africa. One of the best kno ...

. Below is a

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

showing syodontine relationships from a 2012

study of anteosaurians:

References

External links

Palaeos
{{Taxonbar, from=Q263849 Anteosaurs Guadalupian first appearances Guadalupian extinctions Prehistoric therapsid families

Description

Classification

References

Further reading

External links