''Amelanchier alnifolia'', the Saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub with an edible berry-like fruit, native to

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

Description

It is a

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...

or small tree that most often grows to , rarely to , in height. Its growth form spans from suckering and forming

colonies In modern parlance, a colony is a territory subject to a form of foreign rule. Though dominated by the foreign colonizers, colonies remain separate from the administration of the original country of the colonizers, the '' metropolitan state'' ...

to clumped. The

leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

are oval to nearly circular, long and broad, on a

leaf stem In botany, the petiole () is the stalk that attaches the leaf blade to the Plant stem, stem, and is able to twist the leaf to face the sun. This gives a characteristic foliage arrangement to the plant. Outgrowths appearing on each side of the pe ...

, margins toothed mostly above the middle. As with all species in the genus ''

Amelanchier ''Amelanchier'' ( ), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry (or just sarvis), juneberry, saskatoon, sugarplum, wild-plum or chuckley pear,A Digital Flora of Newfoundland and Labrador Vascular Plants/ref> is a g ...

'', the

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...

s are white, with five quite separate petals and five sepals. In ''A. alnifolia'', they are about across, with 20 stamens and five styles, appearing on short

raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...

s of 3–20, somewhat crowded together, blooming from April to July. The fruit is a small purple

pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince. Etymology The word ''pome'' entered English in the late 14th century, and re ...

in diameter, ripening in early summer in the coastal areas and late summer further inland. Resembling blueberries, it has a waxy bloom. Serviceberries are relatively difficult to identify.

Chemistry

Also similar in composition to blueberries, saskatoons have total

polyphenol Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some ...

content of 452 milligrams per 100 grams (average of 'Smoky' and 'Northline'

cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...

s),

flavonol Flavonols are a class of flavonoids that have the 3-hydroxyflavone backbone (IUPAC name : 3-hydroxy-2-phenylchromen-4-one). Their diversity stems from the different positions of the phenolic -OH groups. They are distinct from flavanols (with " ...

s (61 mg) and

anthocyanin Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical com ...

s (178 mg), although others have found the phenolic values to be either lower in the 'Smoky' cultivar or higher.

Quercetin Quercetin is a plant flavonol from the flavonoid group of polyphenols. It is found in many fruits, vegetables, leaves, seeds, and grains; capers, red onions, and kale are common foods containing appreciable amounts of it. It has a bitter flavor ...

cyanidin Cyanidin is a natural organic compound. It is a particular type of anthocyanidin (glycoside version called anthocyanins). It is a pigment found in many red berries including grapes, bilberry, blackberry, blueberry, cherry, chokeberry, cranber ...

, delphinidin,

pelargonidin Pelargonidin is an anthocyanidin, a type of plant pigment producing a characteristic orange color used in food and industrial dyes. Natural occurrences Presence in flowers Pelargonidin can be found in red geraniums (Geraniaceae). It is the ...

petunidin Petunidin (Pt), like Europinidin and Malvidin, is derived from Delphinidin and is an O-methylated anthocyanidin of the 3-hydroxy type. It is a natural organic compound, a dark-red or purple water-soluble pigment found in many redberries includin ...

, peonidin, and

malvidin Malvidin is an O-methylated anthocyanidin, the 3',5'-methoxy derivative of delphinidin. As a primary plant pigment, its glycosides are highly abundant in nature. Natural occurrences Malvidin is responsible for the blue color found in petals ...

were present in saskatoon berries.

Taxonomy

Varieties

The three varieties are:University of Maine
''Amelanchier'' list of taxa
/ref> *''A. a.'' var. ''alnifolia''. Northeastern part of the species' range. *''A. a.'' var. ''pumila'' (Nutt.) A.Nelson.

Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...

Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primar ...

. *''A. a.'' var. ''semiintegrifolia'' (Hook.) C.L.Hitchc. Pacific coastal regions, Alaska to northwestern California.

Etymology

The name ''saskatoon'' derives from the

Cree The Cree ( cr, néhinaw, script=Latn, , etc.; french: link=no, Cri) are a North American Indigenous people. They live primarily in Canada, where they form one of the country's largest First Nations. In Canada, over 350,000 people are Cree o ...

inanimate noun ''misâskwatômina'' ( ''misâskwatômin'' , 'saskatoonberry', ''misâskwatômina'' 'saskatoonberries'). The species name ''alnifolia'' is a feminine

adjective In linguistics, an adjective ( abbreviated ) is a word that generally modifies a noun or noun phrase or describes its referent. Its semantic role is to change information given by the noun. Traditionally, adjectives were considered one of the ...

. It is a compound of the

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

word for "

alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...

", , and the word for "leaf", . Historically, it was also called pigeon berry.

Distribution and habitat

The plant can be found from

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...

across most of

western Canada Western Canada, also referred to as the Western provinces, Canadian West or the Western provinces of Canada, and commonly known within Canada as the West, is a Canadian region that includes the four western provinces just north of the Canada� ...

and in the western and north-central United States. It grows from sea level in the north of the range, up to elevation in

California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...

and in the

. It is a common shrub in the forest

understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...

, as well as canyons.

Ecology

''A. alnifolia'' is susceptible to

cedar-apple rust ''Gymnosporangium juniperi-virginianae'' is a plant pathogen that causes cedar-apple rust. In virtually any location where apples or crabapples (''Malus'') and Eastern red cedar (''Juniperus virginiana'') coexist, cedar apple rust can be a dest ...

, ''entomosporium''

leaf spot A leaf spot is a limited, discoloured, diseased area of a leaf that is caused by fungal, bacterial or viral plant diseases, or by injuries from nematodes, insects, environmental factors, toxicity or herbicides. These discoloured spots or lesions ...

, fireblight, brown rot, '' cytospora'' canker,

powdery mildew Powdery mildew is a fungal disease that affects a wide range of plants. Powdery mildew diseases are caused by many different species of ascomycete fungi in the order Erysiphales. Powdery mildew is one of the easier plant diseases to identify, a ...

, and blackleaf. Problem insects include

aphid Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...

thrips Thrips (order Thysanoptera) are minute (mostly long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Different thrips species feed mostly on plants by puncturing and sucking up the contents, although a few are ...

mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear e ...

s, bud moths, Saskatoon

sawflies Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay ...

, and pear slug sawflies. It is also a larval host to the pale tiger swallowtail, two-tailed swallowtail, and the western tiger swallowtail. The foliage is browsed by deer, elk, rabbits, and livestock. The fruit are eaten by wildlife including birds, squirrels, and bears.

Cultivation

Seedlings are planted with between rows and between plants. An individual bush may bear fruit 30 or more years. Saskatoons are adaptable to most soil types with exception of poorly drained or heavy clay soils lacking organic matter. Shallow soils should be avoided, especially if the water table is high or erratic. Winter hardiness is exceptional, but frost can damage blooms as late as May. Large amounts of sunshine are needed for fruit ripening.

Uses

With a sweet, nutty taste, the fruits have long been eaten by

Indigenous peoples in Canada In Canada, Indigenous groups comprise the First Nations, Inuit and Métis. Although ''Indian'' is a term still commonly used in legal documents, the descriptors ''Indian'' and '' Eskimo'' have fallen into disuse in Canada, and most consider th ...

, fresh or dried. They are well known as an ingredient in

pemmican Pemmican (also pemican in older sources) is a mixture of tallow, dried meat, and sometimes dried berries. A calorie-rich food, it can be used as a key component in prepared meals or eaten raw. Historically, it was an important part of indigenou ...

, a preparation of dried meat to which saskatoon berries are added as flavour and

preservative A preservative is a substance or a chemical that is added to products such as food products, beverages, pharmaceutical drugs, paints, biological samples, cosmetics, wood, and many other products to prevent decomposition by microbial growth or b ...

. They are used in saskatoon berry pie, jam,

wines Wine is an alcoholic drink typically made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different varieties of grapes and strains of yeasts are m ...

cider Cider ( ) is an alcoholic beverage made from the fermented juice of apples. Cider is widely available in the United Kingdom (particularly in the West Country) and the Republic of Ireland. The UK has the world's highest per capita consumption, ...

, beers, and sugar-infused berries similar to dried cranberries used for

cereals A cereal is any grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, germ, and bran. Cereal grain crops are grown in greater quantities and provide more food ...

trail mix Trail mix or scroggin is a type of snack mix, typically a combination of granola, dried fruit, nuts, and sometimes candy, developed as a food to be taken along on hikes. Trail mix is considered a great snack food for hikes, because it is ...

, and

snack foods A snack is a small portion of food generally Eating, eaten between meals. Snacks come in a variety of forms including packaged snack foods and other processed foods, as well as items made from fresh ingredients at home. Traditionally, snack ...

. In 2004, the British

Food Standards Agency The Food Standards Agency is a non-ministerial government department of the Government of the United Kingdom. It is responsible for protecting public health in relation to food in England, Wales and Northern Ireland. It is led by a board ap ...

suspended saskatoon berries from retail sales pending safety testing; the ban eventually was lifted after pressure from the

European Union The European Union (EU) is a supranational union, supranational political union, political and economic union of Member state of the European Union, member states that are located primarily in Europe, Europe. The union has a total area of ...

Nutrition

Saskatoon berries contain significant amounts of total

dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by t ...

riboflavin Riboflavin, also known as vitamin B2, is a vitamin found in food and sold as a dietary supplement. It is essential to the formation of two major coenzymes, flavin mononucleotide and flavin adenine dinucleotide. These coenzymes are involved i ...

and

biotin Biotin (or vitamin B7) is one of the B vitamins. It is involved in a wide range of metabolic processes, both in humans and in other organisms, primarily related to the utilization of fats, carbohydrates, and amino acids. The name ''biotin'', bo ...

, and the

dietary mineral In the context of nutrition, a mineral is a chemical element required as an essential nutrient by organisms to perform functions necessary for life. However, the four major structural elements in the human body by weight (oxygen, hydrogen, carbon ...

s, iron and

manganese Manganese is a chemical element with the Symbol (chemistry), symbol Mn and atomic number 25. It is a hard, brittle, silvery metal, often found in minerals in combination with iron. Manganese is a transition metal with a multifaceted array of ...

, a nutrient profile similar to the content of blueberries.

Culture

The city of

Saskatoon Saskatoon () is the largest city in the Provinces and territories of Canada, Canadian province of Saskatchewan. It straddles a bend in the South Saskatchewan River in the central region of the province. It is located along the Trans-Canada Hig ...

, Saskatchewan, is named after the berry.

References

External links

* * {{Authority control alnifolia Flora of Subarctic America Indigenous cuisine in Canada Plants used in Native American cuisine Trees of the United States Flora of Western Canada Flora of Ontario Flora of the Northwestern United States Flora of the Southwestern United States Flora of the North-Central United States Flora of the South-Central United States