Alstonia

''Alstonia'' is a widespread genus of evergreen trees and shrubs, of the family Apocynaceae. It was named by Robert Brown in 1811, after Charles Alston (1685–1760), professor of botany at Edinburgh from 1716 to 1760.

The type species ''Alstonia scholaris'' (L.) R.Br. was originally named ''Echites scholaris'' by Linnaeus in 1767.

Description

''Alstonia'' consists of about 40–60 species (according to different authors) native to tropical and subtropical Africa, Central America, Southeast Asia, Polynesia and Australia, with most species in the Malesian region.

These trees can grow very large, such as ''Alstonia pneumatophora'', recorded with a height of 60 m and a diameter of more than 2 m. ''Alstonia longifolia'' is the only species growing in Central America (mainly shrubs, but also trees 20 m high).

The leathery, sessile, simple leaves are elliptical, ovate, linear or lanceolate and wedge-shaped at the base. The leaf blade is dorsiventral, medium-sized to large and disposed oppositely or in a whorl and with entire margin. The leaf venation is pinnate, with numerous veins ending in a marginal vein. Phyllotaxy is whorled i.e. two or more leaves arises at a node and form a whorl .

The inflorescence is terminal or axillary, consisting of thyrsiform cymes or compound umbels. The small, more or less fragrant flowers are white, yellow, pink or green and funnel-shaped, growing on a pedicel and subtended by bracts. They consist of 5 petals and 5 sepals, arranged in four whorls. The fertile flowers are hermaphrodite. The gamosepalous green sepals consist of ovate lobes, and are distributed in one whorl. The annular disk is hypogynous. The five gamesepalous petals have oblong or ovate lobes and are disposed in one whorl. The corolla lobes overlapping to the left (such as ''A. rostrata'') or to the right (such as ''A. macrophylla'') in the bud. The ovary has 2 separate follicles with glabrous or ciliate, oblong seeds that develop into deep blue podlike, schizocarp fruit, between 7–40 cm long. The plants contain a milky latex, rich in poisonous alkaloids. The ''Alstonia macrophylla'' is commonly known in Sri Lanka as 'Havari nuga' or the 'wig banyan' because of its distinct flower that looks like a woman's long wig.

''Alstonia'' trees are used in traditional medicine. The bark of the '' Alstonia constricta'' and the '' Alstonia scholaris'' is a source of a remedy against malaria, toothache, rheumatism and snake bites. The latex is used in treating coughs, throat sores and fever.

Many ''Alstonia'' species are harvested for timber, called ''pule'' or ''pulai'' in Indonesia and Malaysia. Trees from the section ''Alstonia'' produce lightweight timber, while those from the sections ''Monuraspermum'' and ''Dissuraspermum'' produce heavy timber.

''Alstonia'' trees are widespread and mostly not endangered. However a few species are very rare, such as ''A. annamensis'', ''A. beatricis'', ''A. breviloba'', ''A. stenophylla'' and ''A. guangxiensis''.

Species

''Alstonia'' has five distinct sections, each a monophyletic group; ''Alstonia'', ''Blaberopus'', ''Tonduzia'', ''Monuraspermum'', ''Dissuraspermum''.

;Accepted species
# '' Alstonia actinophylla'' (A.Cunn.) K.Schum. – milkwood - New Guinea, N Australia
# '' Alstonia angustifolia'' A.DC. - Borneo, W Malaysia, Sumatra
# '' Alstonia angustiloba'' Miq. - Borneo, W Malaysia, Sumatra, Thailand, Java
# '' Alstonia annamensis'' (Monach.) K.Sidiyasa - Cambodia, Vietnam
# '' Alstonia balansae'' Guillaumin - New Caledonia
# '' Alstonia beatricis'' K.Sidiyasa - Waigeo I in E Indonesia
# ''Alstonia boonei'' De Wild. - W + C + E Africa
# '' Alstonia boulindaensis'' Boiteau - New Caledonia
# '' Alstonia breviloba'' K.Sidiyasa - Papua New Guinea
# '' Alstonia congensis'' Engl. - W + C Africa
# '' Alstonia constricta'' F.Muell. – bitterbark, quinine tree, Australian fever bark - E Australia
# '' Alstonia coriacea'' Pancher & S.Moore - New Caledonia
# '' Alstonia costata'' R.Br. - S Pacific
# '' Alstonia curtisii'' King & Gamble - Thailand
# '' Alstonia deplanchei'' Van Heurck & Müll.Arg. - New Caledonia
# '' Alstonia guangxiensis'' D.Fang & X.X.Chen - Guangxi in China
# '' Alstonia iwahigensis'' Elmer - Borneo, Palawan
# '' Alstonia lanceolata'' Van Heurck & Müll.Arg. - New Caledonia
# '' Alstonia lanceolifera'' S.Moore - New Caledonia
# '' Alstonia legouixiae'' Van Heurck & Müll.Arg. - New Caledonia
# '' Alstonia lenormandii'' Van Heurck & Müll.Arg. - New Caledonia
# '' Alstonia longifolia'' (A.DC.) Pichon - Mexico, Central America
# '' Alstonia macrophylla'' Wall. ex G.Don – batino, devil tree - S China, Sri Lanka, SE Asia, New Guinea
# '' Alstonia mairei'' H. Léveillé - S China, N Vietnam
# '' Alstonia muelleriana'' Domin – jackapple, leatherjacket, milky yellowwood - New Guinea, Queensland
# '' Alstonia neriifolia'' D.Don - Nepal, Sikkim, Bhutan
# '' Alstonia odontophora'' Boiteau - New Caledonia
# '' Alstonia parkinsonii'' (M.Gangop. & Chakrab.) Lakra & Chakrab. - Andaman Is.
# '' Alstonia parvifolia'' Merr. - Philippines
# '' Alstonia penangiana'' K.Sidiyasa - Penang Hill in Malaysia
# '' Alstonia pneumatophora'' Backer ex L.G.Den Berger - W Malaysia, Borneo, Sulawesi, Sumatra
# '' Alstonia quaternata'' Van Heurck & Müll.Arg. - New Caledonia
# '' Alstonia rostrata'' C.E.C.Fischer - Yunnan, Indochina, W Malaysia, Sumatra
# '' Alstonia rubiginosa'' K.Sidiyasa - Papua New Guinea
# '' Alstonia rupestris'' Kerr - Thailand
# '' Alstonia scholaris'' (L.) R.Br. – pali-mari, dita bark, bitter bark, milkwood, milky bean, milky pine, white cheesewood, scholar tree, blackboard tree - E + S + SE Asia, Papuasia, N Australia
# '' Alstonia sebusii'' (Van Heurck & Müll.Arg.) Monach. - Yunnan, Bhutan, Assam, N Myanmar
# '' Alstonia spatulata'' Blume – hard milkwood, Siamese balsa - SE Asia, New Guinea
# '' Alstonia spectabilis'' R.Br. – poele bark, jackapple, leatherjacket, milky yellowwood - SE Asia, Papuasia, N Australia
# '' Alstonia sphaerocapitata'' Boiteau - New Caledonia
# '' Alstonia venenata'' R.Br. - S India
# '' Alstonia vieillardii'' Van Heurck & Müll.Arg. - New Caledonia
# '' Alstonia vietnamensis'' D.J.Middleton - Vietnam
# '' Alstonia yunnanensis'' Diels - Yunnan, Guizhou, Guangxi

Gallery

Alstonia scholaris (Saptaparni) in Hyderabad, AP W IMG 1469.jpg, '' Alstonia scholaris''
Alstonia spectabilis Blanco2.379.png, '' Alstonia spectabilis''
Alstonia macrophylla (Batino) in Hyderabad W IMG 7138.jpg, '' Alstonia macrophylla''

Notes

References

*
* Kade Sidiyasa, A., 3, 1992. A monograph of Alstonia (Apocynaceae).
*

{{Taxonbar, from=Q2475725

Apocynaceae genera