Arachnid
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Arachnid
Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Almost all adult arachnids have eight legs, although the front pair of legs in some species has converted to a sensory function, while in other species, different appendages can grow large enough to take on the appearance of extra pairs of legs. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 100,000 named species, of which 47,000 are species of spiders. Morphology Almost all adult arachnids have eight legs, unlike adult inse ...
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Trigonotarbida
The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian ( Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relatives In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 20.5 These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton (tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tube ...
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Spider
Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except for Antarctica, and have become established in nearly every land habitat. , 50,356 spider species in 132 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900. Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segments are fused into two tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindrical pedicel, however, as there is currently neither paleontological nor embryological evidence that spiders ever had ...
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Araneae
Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except for Antarctica, and have become established in nearly every land habitat. , 50,356 spider species in 132 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900. Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segments are fused into two tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindrical pedicel, however, as there is currently neither paleontological nor embryological evidence that spiders ever had a separate ...
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Harvestman
The Opiliones (formerly Phalangida) are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. , over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica. Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their phylogenetic position within the Arachnida is disputed; their closest relatives may be the mites (Acari) or the Novogenuata (the Scorpiones, Pseudoscorpiones, and Solifugae). Althou ...
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Opiliones
The Opiliones (formerly Phalangida) are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. , over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica. Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their phylogenetic position within the Arachnida is disputed; their closest relatives may be the mites (Acari) or the Novogenuata (the Scorpiones, Pseudoscorpiones, and Solifugae). Althou ...
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Ricinulei
Ricinulei is a small order of arachnids. Like most arachnids, they are predatory, eating small arthropods. They occur today in west-central Africa (''Ricinoides'') and the Neotropics (''Cryptocellus'' and ''Pseudocellus'') as far north as Texas. As of 2021, 91 extant species of ricinuleids have been described worldwide, all in the single family Ricinoididae. In older works they are sometimes referred to as Podogona. Due to their obscurity they do not have a proper common name, though in academic literature they are occasionally referred to as hooded tickspiders. In addition to the three living genera, there are fossil species from the upper Carboniferous of Euramerica and the Cretaceous Burmese amber. Description The most important general account of ricinuleid anatomy remains the 1904 monograph by Hans Jacob Hansen and William Sørensen. Useful further studies can be found in, e.g., the work of Pittard and Mitchell, Gerald Legg and L. van der Hammen. Body Ricinulei are typical ...
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Uraraneida
Uraraneida is an extinct order of arachnids, known from fossils of Middle Devonian, Permian and possibly Cretaceous age. Two genera of fossils have been definitively placed in this order: ''Attercopus'' from the Devonian of United States and ''Permarachne'' from the Permian of Russia. In 2018, a third genus ''Chimerarachne'', from the Cretaceous of Myanmar was proposed to belong to this group, but this placement is disputed. When the first fossils were found, they were identified as spiders, but now constitute the Uraraneida, a separate but closely related group. Characteristics The first fossil now placed in the order was found in Gilboa, New York. In 1987, it was initially tentatively placed in the extinct order Trigonotarbida and named ''Gelasinotarbus''? ''fimbriunguis''. Later, partly on the basis of a supposed spinneret (spider), spinneret, it was identified as a spider and named ''Attercopus fimbriunguis''. Further specimens of this species were found, and when examined in ...
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Thelyphonida
Thelyphonida is an arachnid order comprising invertebrates commonly known as whip scorpions or vinegaroons (also spelled vinegarroons and vinegarones). They are often called uropygids in the scientific community based on an alternative name for the order, Uropygi (which may then also include the order Schizomida). The name "whip scorpion" refers to their resemblance to true scorpions and possession of a whiplike tail, and "vinegaroon" refers to their ability when attacked to discharge an offensive, vinegar-smelling liquid, which contains acetic acid. Taxonomy Carl Linnaeus first described a whip scorpion in 1758, although he did not distinguish it from what are now regarded as different kinds of arachnid, calling it ''Phalangium caudatum''. '' Phalangium'' is now used as a name for a genus of harvestmen (Opiliones). In 1802, Pierre André Latreille was the first to use a genus name solely for whip scorpions, namely ''Thelyphonus''. Latreille later explained the name as meaning ...
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Whip Scorpion
Thelyphonida is an arachnid order comprising invertebrates commonly known as whip scorpions or vinegaroons (also spelled vinegarroons and vinegarones). They are often called uropygids in the scientific community based on an alternative name for the order, Uropygi (which may then also include the order Schizomida). The name "whip scorpion" refers to their resemblance to true scorpions and possession of a whiplike tail, and "vinegaroon" refers to their ability when attacked to discharge an offensive, vinegar-smelling liquid, which contains acetic acid. Taxonomy Carl Linnaeus first described a whip scorpion in 1758, although he did not distinguish it from what are now regarded as different kinds of arachnid, calling it ''Phalangium caudatum''. '' Phalangium'' is now used as a name for a genus of harvestmen (Opiliones). In 1802, Pierre André Latreille was the first to use a genus name solely for whip scorpions, namely ''Thelyphonus''. Latreille later explained the name as meaning ...
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Haptopoda
''Plesiosiro'' is an extinct arachnid genus known exclusively from nine specimens from the Upper Carboniferous of Coseley, Staffordshire, United Kingdom. The genus is monotypic, represented only by the species ''Plesiosiro madeleyi'' described by Reginald Innes Pocock in his important 1911 monograph on British Carboniferous arachnids. It is the only known member of the order Haptopoda. The original fossils have been redescribed in detail by Alexander Petrunkevitch in 1949 and Dunlop in 1999. A supposed example from the Coal Measures of Lancashire is a misidentification. Relationships Relationships with other arachnids are obscure. ''Plesiosiro'' means "close to '' Siro''", which is a genus of cyphophthalmid (Cyphophthalmi); the most primitive group of the living harvestmen (Opiliones). These harvestmen do, in some ways, resemble the reconstructed body plan of the haptopods. Revisions have confirmed that Haptopoda should be treated as a separate and independent order. A 2007 stud ...
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Amblypygi
Amblypygi is an ancient order of arachnid chelicerate arthropods also known as African cave-dwelling spiders, whip spiders and tailless whip scorpions (not to be confused with whip scorpions or vinegaroons that belong to the related order Thelyphonida). The name "amblypygid" means "blunt tail", a reference to a lack of the flagellum that is otherwise seen in whip scorpions. Amblypygids possess no silk glands or venomous fangs. They rarely bite if threatened, but can grab fingers with their pedipalps, resulting in thorn-like puncture injuries. As of 2016, 5 families, 17 genera and around 155 species had been discovered and described. They are found in tropical and subtropical regions worldwide; they are mainly found in warm and humid environments and like to stay protected and hidden within leaf litter, caves, or underneath bark. Some species are subterranean; all are nocturnal. Fossilized amblypygids have been found dating back to the Carboniferous period, such as ''Weygoldtina''. ...
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Phalangiotarbi
Phalangiotarbi ( Haase, 1890) is an extinct arachnid order first recorded from the Early Devonian of Germany and most widespread in the Upper Carboniferous coal measures of Europe and North America. The last species are known from the early Permian Rotliegend of Germany. The affinities of phalangiotarbids are obscure, with most authors favouring affinities with Opiliones (harvestmen) and/or Acari (mites and ticks). Phalangiotarbida has been recently (2004) proposed to be sister group to (Palpigradi+Tetrapulmonata): the taxon Megoperculata sensu Shultz (1990). '' Nemastomoides depressus'', described as a harvestman in the family Nemastomoididae, is actually a poorly preserved phalangiotarbid.Dunlop ''in'' Pinto-da-Rocha ''et al.'' 2007: 255 Names The order is also called Phalangiotarbida, the ending -ida originated when Petrunkevitch (1955) tried to standardize the endings of the arachnid orders, which is unnecessary and unwarranted according to the ICZN. Nevertheless, Phalangi ...
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