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Lancelet
The lancelets ( or ), also known as amphioxi (singular: amphioxus ), consist of some 30 to 35 species of "fish-like" benthic filter feeding chordates in the order Amphioxiformes. They are the modern representatives of the subphylum Cephalochordata. Lancelets closely resemble 530-million-year-old ''Pikaia'', fossils of which are known from the Burgess Shale. Zoologists are interested in them because they provide evolutionary insight into the origins of vertebrates. Lancelets contain many organs and organ systems that are closely related to those of modern fish, but in more primitive form. Therefore, they provide a number of examples of possible evolutionary exaptation. For example, the gill-slits of lancelets are used for feeding only, and not for respiration. The circulatory system carries food throughout their body, but does not have red blood cells or hemoglobin for transporting oxygen. Lancelet genomes hold clues about the early evolution of vertebrates: by comparing genes fr ...
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Branchiostoma
''Branchiostoma'' is one of the few living genera of lancelets (order Amphioxiformes). It is the type genus of family Branchiostomatidae. These small vaguely eel- or snake-like animals are close relatives of vertebrates. The scientific name means "gill-mouth", referring to their anatomy – unlike vertebrates, they do not have a true head (with a skull capsule, eyes, nose, a well-developed brain etc.), but merely a mouth adjacent to the gill-slits, with the slightly enlarged anterior end of the dorsal nerve cord above and in front of them. It dislikes the light. Like all lancelets, they are filter feeders that hide in the sediment most of the time. The genus inhabits coastal waters throughout the world. Species * ''Branchiostoma africae'' Hubbs, 1927 * ''Branchiostoma arabiae'' Webb, 1957 * ''Branchiostoma bazarutense'' Gilchrist, 1923 * ''Branchiostoma belcheri'' Gray, 1847 (Belcher's lancelet) * ''Branchiostoma bennetti'' Boschung & Gunter, 1966 (Mud lancelet) * ''Bran ...
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Cephalochordata
A cephalochordate (from Greek: κεφαλή ''kephalé'', "head" and χορδή ''khordé'', "chord") is an animal in the chordate subphylum, Cephalochordata. They are commonly called lancelets. Cephalochordates possess 5 synapomorphies, or primary characteristics, that all chordates have at some point during their larval or adulthood stages. These 5 synapomorphies include a notochord, dorsal hollow nerve cord, endostyle, pharyngeal slits, and a post-anal tail (see chordate for descriptions of each). The fine structure of the cephalochordate notochord is best known for the Bahamas lancelet, ''Asymmetron lucayanum''. Cephalochordates are represented in modern oceans by the Amphioxiformes and are commonly found in warm temperate and tropical seas worldwide. With the presence of a notochord, adult amphioxus are able to swim and tolerate the tides of coastal environments, but they are most likely to be found within the sediment of these communities. Cephalochordates are segmented mar ...
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Pikaia
''Pikaia gracilens'' is an extinct, primitive chordate animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became the "most famous early chordate fossil," or "famously known as the earliest" chordates. It is estimated to have lived during the latter period of the Cambrian explosion. Since it initial discovery, more than a hundred specimens have been recovered. The body structure resembles that of the lancelet and perhaps it swam much like an eel. A notochord and myomeres (segmented blocks of skeletal muscles) span the entire length of the body, and are considered as the defining signatures of chordate characters. Its primitive feature is indicated by the body covering, a cuticle, which is characteristic of invertebrates and some protochordates. The exact phylogenetic position is unclear. Proposed affinities include ...
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Chordates
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cephalochordates. These CSIs prov ...
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Branchiostoma Lanceolatum
''Branchiostoma lanceolatum'', the European lancelet or Mediterranean amphioxus, is a lancelet in the subphylum Cephalochordata. It is a marine invertebrate with a notochord but no backbone and is used as a model organism to study the evolutionary development of vertebrates. Anatomy ''Branchiostoma lanceolatum'' has an elongated body, flattened laterally and pointed at both ends. A stiffening rod of tightly packed cells, the notochord, extends the whole length of the body. Unlike vertebrates, the notochord persists in the adult, in form of simple dorsal neural tube slightly thickened in the anterior part (the cerebral vesicle). Above it is a nerve cord with a single frontal eye. The mouth is on the underside of the body and is surrounded by a tuft of 20 or 30 cirri or slender sensory appendages. The gut runs just below the notochord from the mouth to the anus, in front of the tail. There is a flap-like, vertical fin surrounding the pointed tail. Gas exchange takes place a ...
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Vertebrate
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () (chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do not have ...
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Linda Holland
Linda Zimmerman Holland is a research biologist at Scripps Institution of Oceanography known for her work examining the evolution of vertebrates. Education and career Holland has a B.A. (1962) and an M.A. (1964) from Stanford University. She worked as a research associate at the University of California, San Diego, Scripps Clinic, and Scripps Institution of Oceanography in the period from 1970 until 1998. She earned her Ph.D. from the University of California, San Diego in 2001. She started as a research biologist at Scripps in 1998, and is emeritus as of 2022. Linda Holland has described in detail some of the early obstacles she faced as a woman scientist starting out in academe in the 1960s and 1970s, Research Holland's early research examined anatomical structures in purple sea urchins, and a protein involved in clotting, Von Willebrand factor. She went on to examine reproduction in sea urchins, salps, and amphioxus, known as lancelet. Holland began collecting amphioxus in ...
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Asymmetronidae
''Asymmetron'' is a genus of lancelets belonging to the family Branchiostomatidae. The species of this genus are found in world oceans. Species: *''Asymmetron inferum'' *''Asymmetron lucayanum ''Asymmetron'' is a genus of lancelets belonging to the family Branchiostomatidae. The species of this genus are found in world oceans. Species: *''Asymmetron inferum ''Asymmetron'' is a genus of lancelets belonging to the family Branchiostom ...'' References {{Taxonbar, from=Q9635660 Cephalochordata ...
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Hagfish
Hagfish, of the class Myxini (also known as Hyperotreti) and order Myxiniformes , are eel-shaped, slime-producing marine fish (occasionally called slime eels). They are the only known living animals that have a skull but no vertebral column, although hagfish do have rudimentary vertebrae. Along with lampreys, hagfish are jawless; the two form the sister group to jawed vertebrates, and living hagfish remain similar to hagfish from around 300 million years ago. The classification of hagfish had been controversial. The issue was whether the hagfish was a degenerate type of vertebrate-fish that through evolution had lost its vertebrae (the original scheme) and was most closely related to lampreys, or whether hagfish represent a stage that precedes the evolution of the vertebral column (the alternative scheme) as is the case with lancelets. Recent DNA evidence has supported the original scheme. The original scheme groups hagfish and lampreys together as cyclostomes (or historica ...
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Lampreys
Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like sucking mouth. The common name "lamprey" is probably derived from Latin , which may mean "stone licker" ( "to lick" + "stone"), though the etymology is uncertain. ''Lamprey'' is sometimes seen for the plural form. There are about 38 known extant species of lampreys and five known extinct species. Parasitic carnivorous species are the most well-known, and feed by boring into the flesh of other fish to suck their blood; but only 18 species of lampreys engage in this micropredatory lifestyle. Of the 18 carnivorous species, nine migrate from saltwater to freshwater to breed (some of them also have freshwater populations), and nine live exclusively in freshwater. All non-carnivorous forms are freshwater species. Adults of the non-carnivorous ...
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Branchiostomatidae
Branchiostomatidae is a family of lancelets belonging to the class Leptocardii, order unknown. Genera: * ''Asymmetron'' Andrews, 1893 * '' Branchiostoma'' Costa, 1834 * ''Epigonichthys'' Peters, 1876 * '' Epigonopterus'' Jordan & Gilbert, 1882 * †''Palaeobranchiostoma ''Palaeobranchiostoma hamatotergum'' is an extinct cephalochordate from the Early Permian-aged Whitehill Formation in South Africa.Oelofsen, B. W., and J. C. Loock. "A FOSSIL CEPHALOCHORDATE FROM THE EARLY PERMIAN WHITEHILL FORMATION OF SOUTH-AF ...'' Oelofsen & Loock, 1981 References {{Taxonbar, from1=Q2747865, from2=Q61882392 Cephalochordata ...
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Benthos
Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone.Benthos
from the Census of Antarctic Marine Life website
This community lives in or near marine or freshwater sedimentary environments, from tidal pools along the , out to the continental shelf, and then down to the
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