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Saber-toothed Cat
Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million until about 11,000 years ago. The Machairodontinae contain many of the extinct predators commonly known as "saber-toothed cats", including the famed genus ''Smilodon'', as well as other cats with only minor increases in the size and length of their maxillary canines. The name means "dagger-tooth", from Greek μάχαιρα (''machaira''), sword. Sometimes, other carnivorous mammals with elongated teeth are also called saber-toothed cats, although they do not belong to the felids. Besides the machairodonts, other saber-toothed predators also arose in the nimravids, barbourofelids, machaeroidines, hyaenodonts and even in two groups of metatherians (the thylacosmilid sparassodonts and the deltatheroideans). Evolution Family Felidae ...
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Homotherium
''Homotherium'', also known as the scimitar-toothed cat or scimitar cat, is an extinct genus of machairodontine saber-toothed predator, often termed scimitar-toothed cats, that inhabited North America, South America, Eurasia, and Africa during the Pliocene and Pleistocene epochs (4  mya – 12,000 years ago), existing for approximately . It became extinct in Africa some 1.5 million years ago. The most recent Eurasian remains, recovered from what is now the North Sea, have been dated to around 28,000 years BP. In South America it is only known from a few remains in the northern region (Venezuela), from the mid-Pleistocene. Taxonomy and distribution The name ''Homotherium'' (Greek: (, 'same') and (, 'beast')) was proposed by Emilio Fabrini (1890), without further explanation, for a new subgenus of ''Machairodus'', whose main distinguishing feature was the presence of a large diastema between the two inferior premolars. The lineage of ''Homotherium'' is estimated (based ...
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Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. ...
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Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area and 20% of its land area.Sayre, April Pulley (1999), ''Africa'', Twenty-First Century Books. . With billion people as of , it accounts for about of the world's human population. Africa's population is the youngest amongst all the continents; the median age in 2012 was 19.7, when the worldwide median age was 30.4. Despite a wide range of natural resources, Africa is the least wealthy continent per capita and second-least wealthy by total wealth, behind Oceania. Scholars have attributed this to different factors including geography, climate, tribalism, colonialism, the Cold War, neocolonialism, lack of democracy, and corruption. Despite this low concentration of wealth, recent economic expansion and the large and young population make Afric ...
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Sparassodonta
Sparassodonta (from Greek to tear, rend; and , gen. , ' tooth) is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during th ...
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Thylacosmilidae
Thylacosmilidae is an extinct family of metatherian predators, related to the modern marsupials, which lived in South America between the Miocene and Pliocene epochs. Like other South American mammalian predators that lived prior to the Great American Biotic Interchange, these animals belonged to the order Sparassodonta, which occupied the ecological niche of many eutherian mammals of the order Carnivora from other continents. The family's most notable feature are the elongated, laterally flattened fangs, which is a remarkable evolutionary convergence with other saber-toothed mammals like '' Barbourofelis'' and ''Smilodon''. Taxonomic history The family Thylacosmilidae was originally erected by Riggs in 1933, to accommodate ''Thylacosmilus'', found in the Pliocene Brochero Formation of Argentina. Later, the family was demoted to a subfamily, as Thylacosmilinae, within Borhyaenidae, a group of superficially canid-like sparassodonts, under the assumption that ''Thylacosmilus ...
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Metatheria
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. There are three extant subclasses of mammals, one being metatherians: #monotremes: egg laying mammals like the platypus and the echidna, #metatheria: marsupials, which includes three American orders ( Didelphimorphia, Paucituberculata and Microbiotheria) and four Australasian orders ( Notoryctemorphia, Dasyuromorphia, Peramelemorphia and Diprotodontia), and the # eutherians: placental mammals, consisting of four superorders divided into 21 orders. Metatherians belong to a subgroup of the northern tribosphenic mammal clade or Boreosphenida. They differ from all other mammals in certain morphologies like their dental formula, which includes about five upper and four lower incisors, a canine, three ...
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Hyaenodonta
Hyaenodonta (" hyena teeth") is an extinct order of hypercarnivorous placental pan-carnivoran mammals from mirorder Ferae. Hyaenodonts were important mammalian predators that arose during the early Paleocene in Europe and persisted well into the late Miocene. Characteristics Hyaenodonts are characterized by long skulls, slender jaws, slim bodies and a plantigrade stance. They generally ranged in size from 30 to 140 cm at the shoulder. While '' Simbakubwa kutokaafrika'' may have been up to (surpassing the modern polar bear in size) and ''Hyaenodon gigas'' (the largest species from genus ''Hyaenodon'') was as much as 1.4 m high at the shoulder, 3.0 m long and weighed about 330 kg, most of hyaenodonts were in the 5–15 kg range, equivalent to a mid-sized dog. The anatomy of their skulls show that they had a particularly acute sense of smell, while their teeth were adapted for shearing, rather than crushing. Because of their size range, it is probable that d ...
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Machaeroidinae
Machaeroidinae ("dagger-like") is a subfamily of extinct carnivorous sabre-toothed placental mammals from Asia and North America.Malcolm C. McKenna, Susan K. Bell: ''Classification of Mammals: Above the Species Level'' in Columbia University Press, New York (1997), 631 Seiten. Traditionally classified as hyaenodonts, this group is now classified as a member of the family Oxyaenidae. Classification and phylogeny Taxonomy * Subfamily: †Machaeroidinae (Matthew, 1909) ** Genus: †'' Apataelurus'' (Scott, 1937) *** †''Apataelurus kayi'' (Scott, 1937) *** †''Apataelurus pishigouensis'' (Tong & Lei, 1986) ** Genus: †'' Diegoaelurus'' (Zack, Poust & Wagner, 2022) *** ''Diegoaelurus vanvalkenburghae'' (Zack, Poust & Wagner, 2022) ** Genus: †'' Isphanatherium'' (Lavrov & Averianov, 1998) *** ''Isphanatherium ferganensis'' (Lavrov & Averianov, 1998) ** Genus: †''Machaeroides'' (Matthew, 1909) *** †''Machaeroides eothen'' (Matthew, 1909) *** †''Machaeroides simpsoni'' ( ...
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Barbourofelidae
Barbourofelidae is an extinct family of carnivorans of the suborder Feliformia, sometimes known as false saber-toothed cats, that lived in North America, Eurasia and Africa during the Miocene epoch (16.9—9.0 million years ago) and existed for about . Taxonomy The type genus, ''Barbourofelis'', was originally described by Schultz et al. (1970) and assigned to a new tribe, Barbourofelini, within the felid subfamily Machairodontinae, along with the other sabre-toothed cats. Subsequently, the tribe was reassigned to the Nimravidae by Tedford (1978) and raised to a subfamily by Bryant (1991). However, a number of studies in the early 2000s identified a closer affinity of the barbourofelines to the Felidae than to the Nimravidae and they were reranked as a distinct family by Morlo et al. (2004). Since then the prevailing view has the barbourofelids as the sister group to the Felidae, although this has recently been challenged, following the description of the middle Miocene genus '' ...
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Nimravidae
Nimravidae is an extinct family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America and Eurasia. Not considered to belong to the true cats (family Felidae), the nimravids are generally considered closely related and classified as a distinct family in the suborder Feliformia. Fossils have been dated from the Middle Eocene through the Late Miocene epochs (Bartonian through Tortonian stages, 40.4–7.2 million years ago), spanning about . The barbourofelids, which were formerly classified as a subfamily of the Nimravidae, were reassigned to their own distinct family Barbourofelidae in 2004. However, some recent studies suggest the barbourofelids are a branch of the nimravids, suggesting that this debate might not be settled yet. Morphology and evolution Most nimravids had muscular, low-slung, cat-like bodies, with shorter legs and tails than are typical of cats. Unlike extant Feliformia, the nimravids had a different bone structure ...
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Saber-toothed Predator
A saber-tooth (alternatively spelled sabre-tooth) is any member of various extinct groups of predatory therapsids, predominantly carnivoran mammals, that are characterized by long, curved saber-shaped canine teeth which protruded from the mouth when closed. Saber-toothed mammals have been found almost worldwide from the Eocene epoch to the end of the Pleistocene epoch 42 million years ago ( mya) – 11,000 years ago (kya). One of the best-known genera is the machairodont or "saber-toothed cat" ''Smilodon'', the species of which, especially ''S. fatalis'', are popularly referred to as "saber-toothed tigers", although they are not closely related to tigers ('' Panthera''). Despite some similarities, not all saber-tooths are closely related to saber-toothed cats or felids in-general. Instead, many members are classified into different families of Feliformia, such as Barbourofelidae and Nimravidae; the oxyaenid "creodont" genera ''Machaeroides'' and '' Apataelurus''; and two exti ...
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Maxillary Canine
In human dentistry, the maxillary canine is the tooth located laterally (away from the midline of the face) from both maxillary lateral incisors of the mouth but mesial (toward the midline of the face) from both maxillary first premolars. Both the maxillary and mandibular canines are called the "cornerstone" of the mouth because they are all located three teeth away from the midline, and separate the premolars from the incisors. The location of the canines reflect their dual function as they complement both the premolars and incisors during mastication, commonly known as chewing. Nonetheless, the most common action of the canines is tearing of food. The canines often erupt in the upper gums several millimeters above the gum line. The canine teeth are able to withstand the tremendous lateral pressure caused by chewing. There is a single cusp on canines, and they resemble the prehensile teeth found in carnivorous animals such as the extinct saber-toothed cat. Though relatively t ...
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