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Psocoptera
Psocoptera are a paraphyletic group of insects that are commonly known as booklice, barklice or barkflies. The name Psocoptera has been replaced with Psocodea in recent literature, with the inclusion of the former order Phthiraptera into Psocodea (as part of the suborder Troctomorpha). These insects first appeared in the Permian period, 295–248 million years ago. They are often regarded as the most primitive of the hemipteroids. Their name originates from the Greek word ψῶχος, ''psokhos'' meaning gnawed or rubbed and πτερά, ''ptera'' meaning wings. There are more than 5,500 species in 41 families in three suborders. Many of these species have only been described in recent years. They range in size from 1–10 millimetres (0.04–0.4 in) in length. The species known as booklice received their common name because they are commonly found amongst old books—they feed upon the paste used in binding. The barklice are found on trees, feeding on algae and ...
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Paraneoptera
Paraneoptera or Acercaria is a superorder of insects which includes lice (bark lice and true lice), thrips, and hemipterans, the true bugs. It also includes the extinct order Permopsocida, known from fossils dating from the Early Permian to the mid-Cretaceous. All of the insects classified here exhibit various “reductions” or “simplifications” from the primitive body-plan found in typical polyneopterans. Cerci, for example, are entirely absent in all living paraneopterans (Acercaria meaning without cerci). Other “reductions” occur in wing venation, in the number of tarsal segments (no more than three), only four Malpighian tubules, and only one complex of abdominal ganglia. The mouthparts of the Paraneoptera reflect diverse feeding habits. Basal groups are microbial surface feeders, whereas more advanced groups feed on plant or animal fluids. Phylogeny Paraneoptera consists of Psocodea (lice), along with their sister clade, the monophyletic grouping Condylognat ...
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Psocodea
Psocodea is a taxonomic group of insects comprising the bark lice, book lice and parasitic lice. It was formerly considered a superorder, but is now generally considered by entomologists as an order. Despite the greatly differing appearance of parasitic lice (Phthiraptera), they are believed to have evolved from within the former order Psocoptera, which contained the bark lice and book lice, now found to be paraphyletic. They are often regarded as the most primitive of the hemipteroids. Psocodea contains around 11,000 species, divided among four suborders and more than 70 families. They range in size from 1–10 millimetres (0.04–0.4 in) in length. The species known as booklice received their common name because they are commonly found amongst old books—they feed upon the paste used in binding. The barklice are found on trees, feeding on algae and lichen. Anatomy and biology Psocids are small, scavenging insects with a relatively generalized body plan. They feed ...
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Psocomorpha
Psocomorpha is a suborder of barklice, booklice, and parasitic lice in the order Psocodea (formerly Psocoptera). There are more than 20 families and 5,300 described species in Psocomorpha. Phylogeny The below cladogram of Psocodea shows the position of Psocomorpha: Fossil record The oldest fossil record of the suborder is '' Burmesopsocus lienhardi'' from the mid-Cretaceous (early Cenomanian) Burmese amber, which is not assigned to any family. Classification Psocomorpha contains 6 infraorders and 26 families: * Archipsocetae ** Archipsocidae Pearman, 1936 (ancient barklice) *Caeciliusetae ** Amphipsocidae Pearman, 1936 (hairy-winged barklice) ** Asiopsocidae Mockford & Garcia Aldrete, 1976 ** Caeciliusidae Mockford, 2000 (lizard barklice) ** Dasydemellidae Mockford, 1978 (shaggy psocids) ** Paracaeciliidae Mockford, 1989 ** Stenopsocidae Pearman, 1936 (narrow barklice) * Epipsocetae ** Cladiopsocidae Smithers, 1972 ** Dolabellopsocidae Eertmoed, 1973 ** Epipsocidae Pear ...
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Troctomorpha
Troctomorpha is one of the three major suborders of Psocodea (formerly Psocoptera)(barklice, booklice, and parasitic lice), alongside Psocomorpha and Trogiomorpha. There are more than 30 families and 5,800 described species in Troctomorpha. The order includes parasitic lice, which are most closely related to the booklice family Liposcelididae. Cladogram Cladogram showing the position of Troctomorpha within Psocodea: Fossil record The oldest record of the suborder is suggested to be '' Paramesopsocus adibi,'' known from the Late Jurassic Karabastau Formation of Kazakhstan. Classification Troctomorpha contains the following subgroups: * Infraorder: Amphientometae ** Family: Amphientomidae Enderlein, 1903 (tropical barklice) ** Family: Compsocidae Mockford, 1967 ** Family: Electrentomidae Enderlein, 1911 ** Family: Manicapsocidae Mockford, 1967 ** Family: Musapsocidae Mockford, 1967 ** Family: Protroctopsocidae Smithers, 1972 ** Family: Troctopsocidae Mockford, 19 ...
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Phthiraptera
Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a result of developments in phylogenetic research. Lice are obligate parasites, living externally on warm-blooded hosts which include every species of bird and mammal, except for monotremes, pangolins, and bats. Lice are vectors of diseases such as typhus. Chewing lice live among the hairs or feathers of their host and feed on skin and debris, while sucking lice pierce the host's skin and feed on blood and other secretions. They usually spend their whole life on a single host, cementing their eggs, called nits, to hairs or feathers. The eggs hatch into nymphs, which moult three times before becoming fully grown, a process that takes about four weeks. Genetic evidence indicates that lice are a highly modified lineage of Psocoptera (now called Psoco ...
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Phthiraptera
Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a result of developments in phylogenetic research. Lice are obligate parasites, living externally on warm-blooded hosts which include every species of bird and mammal, except for monotremes, pangolins, and bats. Lice are vectors of diseases such as typhus. Chewing lice live among the hairs or feathers of their host and feed on skin and debris, while sucking lice pierce the host's skin and feed on blood and other secretions. They usually spend their whole life on a single host, cementing their eggs, called nits, to hairs or feathers. The eggs hatch into nymphs, which moult three times before becoming fully grown, a process that takes about four weeks. Genetic evidence indicates that lice are a highly modified lineage of Psocoptera (now called Psoco ...
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Trogiomorpha
Trogiomorpha is one of the three major suborders of barklice, booklice, and parasitic lice in the order Psocodea (formerly Psocoptera), alongside Troctomorpha and Psocomorpha. There are about 8 families and more than 430 described species in Trogiomorpha. Trogiomorpha is widely agreed to be the earliest diverging of the three suborders, and retains the most primitive characteristics. Internal phylogeny Here is a cladogram showing the phylogeny of Psocodea, showing the position of Trogiomorpha: Families Psocomorpha contains 3 infraorders and 5 extant (living) families, as well as three identified extinct families: * Atropetae ** Archaeatropidae Baz & Ortuño, 2000 ** Empheriidae Baz & Ortuño, 2000 ** Lepidopsocidae Enderlein, 1903 (scaly-winged barklice) ** Psoquillidae Lienhard & Smithers, 2002 (bird nest barklice) ** Trogiidae Roesler, 1944 (granary booklice) * Psyllipsocetae ** Psyllipsocidae Lienhard & Smithers, 2002 (cave barklice) * Prionoglaridetae (paraphyletic ...
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Insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch f ...
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Insect Wing
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Tho ...
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Insect Wing
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Tho ...
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Mandible (arthropod)
250px, The mandibles of a bull ant The mandible (from la, mandibula or mandĭbŭ-lum, a jaw) of an arthropod is a pair of mouthparts used either for biting or cutting and holding food. Mandibles are often simply called jaws. Mandibles are present in the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects etc.). These groups make up the clade Mandibulata, which is currently believed to be the sister group to the rest of arthropods, the clade Arachnomorpha (Chelicerata and Trilobita). Unlike the chelicerae of arachnids, mandibles can often be used to chew food. Mandibulates also differ by having antennae, and also by having three distinct body regions: head, thorax and abdomen. (The cephalothorax (or prosoma) of chelicerates is a fusion of head and thorax.) Insects Insect mandibles are as diverse in form as their food. For instance, grasshoppers and many other plant-eating insects have sharp-edged mandibles that move side to side. Mo ...
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Insect Mouthparts
Insects have mouthparts that may vary greatly across insect species, as they are adapted to particular modes of feeding. The earliest insects had chewing mouthparts. Most specialisation of mouthparts are for piercing and sucking, and this mode of feeding has evolved a number of times idependently. For example, mosquitoes and aphids (which are true bugs) both pierce and suck, however female mosquitoes feed on animal blood whereas aphids feed on plant fluids. Evolution Like most external features of arthropods, the mouthparts of Hexapoda are highly derived. Insect mouthparts show a multitude of different functional mechanisms across the wide diversity of insect species. It is common for significant homology to be conserved, with matching structures forming from matching primordia, and having the same evolutionary origin. However, even if structures are almost physically and functionally identical, they may not be homologous; their analogous functions and appearance might be the ...
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