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Parenthesome
Within the cells of some members of basidiomycetes fungi are found microscopic structures called parenthesomes or septal pore caps. They are shaped like parentheses and found on either side of pores in the dolipore septum which separates cells within a hypha A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or .... Their function has not been established, and their composition has not been fully elucidated. The variations in their appearance are useful in distinguishing individual species. Generally, they are barrel shaped, with an endoplasmic reticulum covering. See also * Pit connection References * * Organelles Mycology Fungal morphology and anatomy {{Basidiomycota-stub ...
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Dolipore Septum
Dolipore septa are specialized dividing walls between cells (septa) found in almost all species of fungi in the phylum Basidiomycota. Unlike most fungal septa, they have a barrel-shaped swelling around their central pore, which is about 0.1–0.2 µm wide. This structure is typically capped at either end by specialized membranes, called " parenthesomes" (after their parenthesis-like appearance under a microscope) or simply "pore caps". Dolipore septa vary significantly between monokaryotic and dikaryotic hyphae, which form at different points in basidiomycete life cycles. In monokaryotic but not dikaryotic hyphae, the parenthesomes are continuous with the endoplasmic reticulum, and the septal walls are constructed from different material than the cell walls. All dolipore septa can allow cytoplasm, and sometimes mitochondria, to flow through their pores; those in monokaryotic hyphae have perforated parenthesomes, which allow cell nuclei The cell nucleus (pl. nuclei; ...
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Basidiomycetes
Basidiomycota () is one of two large divisions that, together with the Ascomycota, constitute the subkingdom Dikarya (often referred to as the "higher fungi") within the kingdom Fungi. Members are known as basidiomycetes. More specifically, Basidiomycota includes these groups: mushrooms, puffballs, stinkhorns, bracket fungi, other polypores, jelly fungi, boletes, chanterelles, earth stars, smuts, bunts, rusts, mirror yeasts, and ''Cryptococcus'', the human pathogenic yeast. Basidiomycota are filamentous fungi composed of hyphae (except for basidiomycota-yeast) and reproduce sexually via the formation of specialized club-shaped end cells called basidia that normally bear external meiospores (usually four). These specialized spores are called basidiospores. However, some Basidiomycota are obligate asexual reproducers. Basidiomycota that reproduce asexually (discussed below) can typically be recognized as members of this division by gross similarity to others, by the forma ...
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Fungus
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota ...
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Bracket
A bracket is either of two tall fore- or back-facing punctuation marks commonly used to isolate a segment of text or data from its surroundings. Typically deployed in symmetric pairs, an individual bracket may be identified as a 'left' or 'right' bracket or, alternatively, an "opening bracket" or "closing bracket", respectively, depending on the directionality of the context. Specific forms of the mark include parentheses (also called "rounded brackets"), square brackets, curly brackets (also called 'braces'), and angle brackets (also called 'chevrons'), as well as various less common pairs of symbols. As well as signifying the overall class of punctuation, the word "bracket" is commonly used to refer to a specific form of bracket, which varies from region to region. In most English-speaking countries, an unqualified word "bracket" refers to the parenthesis (round bracket); in the United States, the square bracket. Various forms of brackets are used in mathematics, with s ...
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Hypha
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 µm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membrane. T ...
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Pit Connection
In algal anatomy, a pit connection is a hole in the septum between two algal cells, and is found only in the red algae − specifically, all orders except the Porphyridiales and haploid Bangiales. They are often stoppered with proteinaceous "pit plugs". By contrast, many fungi (only ascomycetes and basidiomycetes, as most other groups lack septa) contain septal pores − an unrelated phenomenon. Characteristics A sieve-like membrane may cover the pit in living algae, but in the majority of algae a plug forms, they likely limit the transfer of metabolites between neighbouring cells. Formation Primary pit connections are formed between cells in the same filament, derived from the same parent cell by its division. Such connections are always single, and usually circular; this is a result of their method of formation. The septum is formed as the walls of a filament grow inwards, dividing the cell; this results in a hole in the middle of the tube where the walls don't quite merge. T ...
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Organelles
In cell biology, an organelle is a specialized subunit, usually within a cell, that has a specific function. The name ''organelle'' comes from the idea that these structures are parts of cells, as organs are to the body, hence ''organelle,'' the suffix ''-elle'' being a diminutive. Organelles are either separately enclosed within their own lipid bilayers (also called membrane-bound organelles) or are spatially distinct functional units without a surrounding lipid bilayer (non-membrane bound organelles). Although most organelles are functional units within cells, some function units that extend outside of cells are often termed organelles, such as cilia, the flagellum and archaellum, and the trichocyst. Organelles are identified by microscopy, and can also be purified by cell fractionation. There are many types of organelles, particularly in eukaryotic cells. They include structures that make up the endomembrane system (such as the nuclear envelope, endoplasmic reticulum, an ...
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Mycology
Mycology is the branch of biology concerned with the study of fungi, including their genetic and biochemical properties, their taxonomy and their use to humans, including as a source for tinder, traditional medicine, food, and entheogens, as well as their dangers, such as toxicity or infection. A biologist specializing in mycology is called a mycologist. Mycology branches into the field of phytopathology, the study of plant diseases, and the two disciplines remain closely related because the vast majority of plant pathogens are fungi. Overview Historically, mycology was a branch of botany because, although fungi are evolutionarily more closely related to animal Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage ...s than to plants, this was not recognized until a few ...
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