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Panpulmonata
Panpulmonata is a taxonomic clade of snails and slugs in the clade Heterobranchia within the clade Euthyneura. Panpulmonata was established as a new taxon by Jörger et al. in October 2010. The older name "Pulmonata" referred to a group of gastropods which were considered to be "air-breathers". This meaning certainly does not apply to the panpulmonate groups Acochlidia, Sacoglossa and Pyramidelloidea, and also was inaccurate when applied to some of the more traditional pulmonate taxa such as Siphonarioidea or Hygrophila, most members of which lack permanently air-filled lungs. However, the term Panpulmonata was chosen by Jörger et al. (2010) to provide some continuity in the terminology. Panpulmonata consists of following taxa: * Siphonarioidea * Sacoglossa * Glacidorboidea * Amphiboloidea * Pyramidelloidea * Hygrophila * Acochlidiacea (mentioned as Acochlidia) * Eupulmonata: Stylommatophora, Systellommatophora, Ellobioidea, Otinoidea, Trimusculoidea. Cladogram T ...
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Hygrophila (Gastropoda)
Hygrophila is a taxonomic superorder of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks within the clade Panpulmonata. MolluscaBase eds. (2020). MolluscaBase. Hygrophila. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=382243 on 2020-12-14 The families in this clade are basically air-breathing freshwater snails including freshwater limpets. The three families with the greatest number of species are the Lymnaeidae (pond snails), the Planorbidae (ramshorn snails) and the Physidae (pouch or bubble snails). These are found in ponds, creeks, ditches, and shallow lakes nearly worldwide. The snails in this clade have their eyes located at the base of their tentacles, rather than at the tips, as in the true land snails Stylommatophora. They have shells that are thin, translucent, and relatively colorless, and they lack an operculum. Taxonomy 1997 taxonomy In the older taxonomy of the Gastro ...
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Siphonarioidea
Siphonarioidea is a taxonomic superfamily of air-breathing sea snails or false limpets, marine gastropod mollusks in the clade Panpulmonata. Distribution The Siphonarioidea are broadly distributed across the globe, however they are known to be the most abundant and species diverse in the warmer temperate to subtropical regions. More specifically, they are found within the southern hemisphere of the Indo-Pacific region. They generally inhabit marine benthic regions, specifically the subtidal zones. Description Siphonarioidea are almost entirely marine organisms, as they have been found to have evolved into despite the terrestrial ecosystems inhabited by their ancestors. They inhabit the subtidal zones of marine ecosystems, and resemble true limpets with their non coiled, conical shells. Siphonarioidea are frequently referred to as "false limpets" due to some distinct differences despite their resemblance to these Patellogastropoda. For example, Siphonariid adults' dorso-v ...
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Pyramidelloidea
Pyramidelloidea is a superfamily of mostly very small sea snails, marine gastropod mollusks and micromollusks within the clade Panpulmonata. This is a voluminous taxon: above the species level close to 400 named taxa are referred to this gastropod superfamily. Pyramidelloidea has both fossil and recent members. They live as ectoparasites on bivalve molluscs and polychaete worms, and have a sharp, piercing stylet instead of a radula. Distribution This taxon is found worldwide. Taxonomy 1999 taxonomy Taxonomy by Schander, Van Aartsen & Corgan (1999): *Superfamily Pyramidelloidea Gray, 1840 **Family Amathinidae Ponder, 1987 **Family Ebalidae Warén, 1994 - synonym: Anisocyclidae van Aartsen, 1995 **Family Odostomiidae Pelseneer, 1928 ***Subfamily Odostomiinae Pelseneer, 1928 ***Subfamily Chrysallidinae Saurin, 1958 ***Subfamily Odostomellinae Saurin, 1958 ***Subfamily Cyclostremellinae Moore, 1966 **Family Pyramidellidae J. E. Gray, 1840 ***Subfamily Pyramide ...
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Heterobranchia
Heterobranchia, the ''heterobranchs'' (meaning "different-gilled snails"), is a taxonomic clade of snails and slugs, which includes marine, aquatic and terrestrial gastropod mollusks. Heterobranchia is one of the main clades of gastropods. Currently Heterobranchia comprises three informal groups: the lower heterobranchs, the opisthobranchs and the pulmonates.Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdes A. & Warén A. 2005. ''Classification and nomenclator of gastropod families''. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. . . 397 pp. http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278 Diversity The three subdivisions of this large clade are quite diverse: * The Lower Heterobranchia includes shelled marine and freshwater species. * Opisthobranchia are almost all marine species, some shelled and some not. The internal organs of the opisthobranchs have undergone detorsion (unwinding of the vi ...
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Euthyneura
Euthyneura is a taxonomic infraclass of snails and slugs, which includes species exclusively from marine, aquatic and terrestrial gastropod mollusks in the clade Heterobranchia. Euthyneura are characterised by several autapomorphies, but are named for euthyneury. They are considered to be the most successful and diverse group of Gastropoda. Within this taxon, the Gastropoda have reached their peak in species richness and ecological diversity. This obvious evolutionary success can probably be attributed to several factors. Marine Opisthobranchia, e.g., have evolved several clades specialised on less used food resources such as sponges or cnidarians. A key innovation in the evolution of Pulmonata was the colonization of freshwater and terrestrial habitats.Klussmann-Kolb A., Dinapoli A., Kuhn K., Streit B. & Albrecht C. (2008). "From sea to land and beyond – New insights into the evolution of euthyneuran Gastropoda (Mollusca)". '' BMC Evolutionary Biology'' 2008, 8: 57. . ...
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Sacoglossa
Sacoglossa, commonly known as the sacoglossans or the "solar-powered sea slugs", are a superorder of small sea slugs and sea snails, marine gastropod mollusks that belong to the clade Heterobranchia. Sacoglossans live by ingesting the cellular contents of algae, hence they are sometimes called "sap-sucking sea slugs". Some sacoglossans simply digest the fluid which they suck from the algae, but in some other species, the slugs sequester and use within their own tissues living chloroplasts from the algae they eat, a very unusual phenomenon known as kleptoplasty, for the "stolen" plastids. This earns them the title of the "solar-powered sea slugs", and makes them unique among metazoan organisms, for otherwise kleptoplasty is known only among single-celled protists. The Sacoglossa are divided into two clades - the shelled families (Oxynoacea) and the shell-less families (Plakobranchacea). The four families of shelled species are: Cylindrobullidae, Volvatellidae, Oxynoidae, and J ...
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Acochlidia
Acochlidiacea, common name acochlidians, are a taxonomic clade of very unusual sea snails and sea and freshwater slugs, aquatic gastropod mollusks within the large clade Heterobranchia. Acochlidia is a variant spelling. Description These are mostly very small animals, without a shell or gills, distinguished by the visceral mass being sharply set off from the rest of the body. Being a small group with only 30 species worldwide known in 2010, and 32 species described in 2011, and 33 in 2012 (+9 undescribed ''Pontohedyle'' species), these slugs are morphologically and biologically highly aberrant and diverse, comprising a series of unusual characters (e.g. secondary gonochorism, lack of copulatory organs, asymmetric radulae). Most acochlidians live interstitially in marine sands, while some have conquered limnic systems (uniquely within opisthobranch gastropods). Taxonomy Nils Hjalmar Odhner established this taxon as a family in 1937, when he created the families Microhe ...
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Amphiboloidea
Amphiboloidea is a taxonomic superfamily of air-breathing land snails. Distribution Amphibolids are found in Indo-Pacific intertidal mangrove, saltmarsh and estuarine mudflat habitats. Taxonomy 2005 taxonomy According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), it is a superfamily in the informal group Basommatophora, within the Pulmonata. This superfamily has contained only one family, the Amphibolidae. 2007 taxonomy Golding et al. (2007)Golding R. E., Ponder W. F. & Byrne M. (2007). "Taxonomy and anatomy of Amphiboloidea (Gastropoda: Heterobranchia: Archaeopulmonata)". ''Zootaxa'' 1476: 1-50abstract have established new families: * Maningrididae Golding, Ponder & Byrne, 2007 - with the only species ''Maningrida arnhemensis'' * Phallomedusidae Golding, Ponder & Byrne, 2007 2010 taxonomy Basommatophora ( Siphonarioidea and Amphiboloidea and Hygrophila) have been found polyphyletic and so Jörger et al. (2010)Jörger K. M., Stöger I., Kano Y., Fuku ...
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Acochlidiacea
Acochlidiacea, common name acochlidians, are a taxonomic clade of very unusual sea snails and sea and freshwater slugs, aquatic gastropod mollusks within the large clade Heterobranchia. Acochlidia is a variant spelling. Description These are mostly very small animals, without a shell or gills, distinguished by the visceral mass being sharply set off from the rest of the body. Being a small group with only 30 species worldwide known in 2010, and 32 species described in 2011, and 33 in 2012 (+9 undescribed '' Pontohedyle'' species), these slugs are morphologically and biologically highly aberrant and diverse, comprising a series of unusual characters (e.g. secondary gonochorism, lack of copulatory organs, asymmetric radulae). Most acochlidians live interstitially in marine sands, while some have conquered limnic systems (uniquely within opisthobranch gastropods). Taxonomy Nils Hjalmar Odhner established this taxon as a family in 1937, when he created the families Microhedy ...
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Pulmonata
Pulmonata or pulmonates, is an informal group (previously an Order (biology), order, and before that a Class (biology), subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a Respiratory system of gastropods#Pulmonates, pallial lung instead of a gill, or gills. The group includes many land and freshwater families, and several marine families. The taxon Pulmonata as traditionally defined was found to be polyphyletic in a molecular study per Jörger ''et al.'', dating from 2010. Pulmonata are known from the Carboniferous Period to the present. Pulmonates have a single Atrium (heart), atrium and kidney, and a concentrated, symmetrical, nervous system. The mantle cavity is located on the right side of the body, and lacks gills, instead being converted into a blood vessel, vascularised lung. Most species have a shell, but no operculum (gastropod), operculum, although the group does also include several shell-less slugs. Pulmonates are hermap ...
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Euopisthobranchia
Euopisthobranchia is a taxonomic clade of snails and slugs in the clade Heterobranchia within the clade Euthyneura. This clade was established as a new taxon by Jörger et al. in October 2010. Euopisthobranchia is a monophyletic portion of the Opisthobranchia as that taxon was traditionally defined but is not a replacement name for that group as several marine opisthobranch orders including Nudibranchia, Sacoglossa and Acochlidiacea are not included. Euopisthobranchia consist of the following taxa: * Umbraculoidea * Anaspidea * Runcinacea * Pteropoda * Cephalaspidea s.s. Gizzard Previous studies discussed the gizzard (i.e. a muscular oesophageal crop lined with cuticula) with gizzard plates as homologous apomorphic structures supporting a clade composed of Cephalaspidea s.s., Pteropoda and Anaspidea. A gizzard with gizzard plates probably originated in herbivorous taxa in which it worked like a grinding mill, thus might be secondarily reduced in carnivorous groups within ...
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Glacidorboidea
The Glacidorbidae is a taxonomic family of freshwater snails. Taxonomy 2005 taxonomy Glacidorbidae is the only family in the superfamily Glacidorboidea. Glacidorboidea has been classified within the informal group Lower Heterobranchia in the taxonomy of Bouchet & Rocroi (2005). 2010 taxonomy Jörger et al. (2010) redefined the major groups within the Heterobranchia; they moved Glacidorboidea to Panpulmonata.Jörger K. M., Stöger I., Kano Y., Fukuda H., Knebelsberger T. & Schrödl M. (2010). "On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia". ''BMC Evolutionary Biology ''BMC Ecology and Evolution'' (since January 2021), previously ''BMC Evolutionary Biology'' (2001–2020), is a peer-reviewed open access scientific journal covering all fields of evolutionary biology, including phylogenetics and palaeontology ...'' 10: 323. . Genera Genera within the family Glacidorbidae include:Ponder, ...
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