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Ophiacodontidae
Ophiacodontidae is an extinct family of early eupelycosaurs from the Carboniferous and Permian. '' Archaeothyris'', and ''Clepsydrops'' were among the earliest ophiacodontids, appearing in the Late Carboniferous. Ophiacodontids are among the most basal synapsids, an offshoot of the lineage which includes therapsids and their descendants, the mammals. The group became extinct by the Middle Permian, replaced by anomodonts, theriodonts, and the diapsid reptiles. Characteristics The lifestyle of ophiacodonts has long been controversial. Some studies suggested that they were semi-aquatic, and some even suggested a fairly aquatic lifestyle, but a recent study based on a quantitative inference model suggested that both ''Clepsydrops'' and '' Ophiacodon'' were terrestrial. Vertebral morphometric data also support, though ambiguously, a rather terrestrial lifestyle for '' Ophiacodon'', which could reach a length of . '' Archaeothyris'' may also have been terrestrial, but no detailed s ...
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Synapsid
Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptiles and birds. The group includes mammals and every animal more closely related to mammals than to sauropsids. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name. The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals. Traditionally, non-mammalian synapsids were believed to have evolved from reptiles, and therefore described as mammal-like reptiles in classical systematics, and primitive synapsids were also referred to as pelycosaurs, or pelycosaur- ...
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Eupelycosauria
Eupelycosauria is a large clade of animals characterized by the unique shape of their skull, encompassing all mammals and their closest extinct relatives. They first appeared 308million years ago during the Early Pennsylvanian epoch, with the fossils of '' Echinerpeton'' and perhaps an even earlier genus, '' Protoclepsydrops'', representing just one of the many stages in the evolution of mammals,Kemp. T.S., 1982, ''Mammal-like Reptiles and the Origin of Mammals''. Academic Press, New York in contrast to their earlier amniote ancestors. Eupelycosaurs are synapsids, animals whose skull has a single opening behind the eye. They are distinguished from the Caseasaurian synapsids by having a long, narrow supratemporal bone (instead of one that is as wide as it is long) and a frontal bone with a wider connection to the upper margin of the orbit. Laurin, M. and Reisz, R. R., 1997Autapomorphies of the main clades of synapsids- Tree of Life Web Project The only living descendants of ...
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Baldwinonus
''Baldwinonus'' is an extinct genus of basal synapsids from the Early Permian. The type species is ''Baldwinonus trux'', named in 1940 from the Cutler Formation of New Mexico. A second species, ''Baldwinonus dunkardensis'', was named in 1952 from Ohio. ''Baldwinonus'' was first classified in the family Eothyrididae, but the group has since been recognized as a wastebasket taxon for many early synapsids.R. R. Reisz. 1986. Pelycosauria. Handbuch der Paläoherpetologie / Encyclopedia of Paleoherpetology 17A:1-102 More recently, ''Baldwinonus'' has been placed in the family Ophiacodontidae. Its phylogenetic relationship to other early synapsids remains poorly understood because it is only known from a few fragments of bone. Description ''Baldwinonus trux'' is known from a fragment of a right maxilla or upper jaw bone, part of a quadrate bone, and several vertebrae. The maxilla contains 25 tooth sockets, some with teeth. There are sockets for five precaniniform teeth at the front of ...
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Archaeothyris
''Archaeothyris'' is an extinct genus of ophiacodontid synapsid that lived during the Late Carboniferous and is known from Nova Scotia. Dated to 306 million years ago, ''Archaeothyris'', along with a more poorly known synapsid called '' Echinerpeton'', are the oldest undisputed synapsids known. The name means ''ancient window'' (Greek), and refers to the opening in the skull, the temporal fenestra, which indicates this is an early synapsid. '' Protoclepsydrops'' also from Nova Scotia is slightly older but is known by very fragmentary materials. Description ''Archaeothyris'' was more advanced than the early sauropsids, having strong jaws that could open wider than those of the early reptiles. While its sharp teeth were all of the same size & shape, it did possess a pair of enlarged canines, suggesting that it was a carnivore. Archaeothyris' legs were articulated laterally at its pelvis and shoulders, which gave it a sprawling stance. The first toe is smaller than the second. ...
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Protoclepsydrops
''Protoclepsydrops'' is an extinct genus of early synapsids, found in Joggins, Nova Scotia. The name means 'first ''Clepsydrops, and refers to it being the predecessor of the other early synapsid ''Clepsydrops''. Description Like ''Archaeothyris'', ''Protoclepsydrops'' resembled a modern lizard in superficial appearance. However, ''Protoclepsydrops'' had primitive vertebrae with tiny neural processes typical of their amniote ancestors. ''Protoclepsydrops'' is known from a few vertebrae and some humeri.Reisz, R., 1972Pelycosaurian reptiles from the middle Pennsylvanian of North America. Harvard University Classification Its skeletal remains indicate that it may have been more closely related to synapsids than to sauropsids, making it a possible stem-mammal. If so, it is the oldest synapsid known, though its status is unconfirmed because its remains are too fragmentary. ''Protoclepsydrops'' lived slightly earlier than ''Archaeothyris''.Benton M.J. and Donoghue P.C.J. 2006. Palaeon ...
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Ophiacodon Mirus
''Ophiacodon'' (meaning "snake tooth") is an extinct genus of synapsid belonging to the family Ophiacodontidae that lived from the Late Carboniferous to the Early Permian in North America and Europe. The genus was named along with its type species ''O. mirus'' by paleontologist Othniel Charles Marsh in 1878 and currently includes five other species. As an ophiacodontid, ''Ophiacodon'' is one of the most basal synapsids and is close to the evolutionary line leading to mammals. Description ''Ophiacodon'' has a large skull with a deep snout. It has the longest skull of any early synapsid, reaching up to in one specimen. The jaws are lined with many small teeth. It was larger than most other tetrapods (four-limbed vertebrates) of its time, ranging from in length and in weight. Specimens of ''Ophiacodon'' vary greatly in size. These differences in size were once used to distinguish species, but are now recognized as ontogenetic variations related to the ages of individuals. S ...
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Varanosaurus
''Varanosaurus'' (' monitor lizard') is an extinct genus of early pelycosaur synapsid that lived during the Kungurian. Description As its name implies, ''Varanosaurus'' may have looked superficially similar to present-day monitor lizards, though not related at all. ''Varanosaurus'' had a flattened, elongated skull and a pointed snout with a row of sharp teeth, including two pairs of conspicuous pseudocanines, implying that it was an active predator. ''Varanosaurus'' probably lived in swamps, competing with the larger '' Ophiacodon'' for food. Classification Below is a cladogram modified from the analysis of Benson (2012): See also * List of pelycosaurs This list of pelycosaurs is an attempt to create a comprehensive listing of all genera that have ever been included in the synapsida excluding therapsida and purely vernacular terms. The list includes all commonly accepted genera, but also genera t ... References Further reading *Benes, Josef. Prehistoric Animals and P ...
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Stereorhachis
''Stereorhachis'' is an extinct genus of non-mammalian synapsids from the Late Carboniferous Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ... of France. See also * List of pelycosaurs References * Sumida, S. J. & Martin, K. L. M. (eds.). ''Amniote Origins - Completing the Transition to Land''. Academic Press, 1997. 510 p. * Carroll, R. L. ''Vertebrate Paleontology and Evolution''. W. H Freeman Company, 1988. Ophiacodontids Carboniferous synapsids Fossils of France Fossil taxa described in 1880 Prehistoric synapsid genera {{paleo-Synapsid-stub ...
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Ophiacodon
''Ophiacodon'' (meaning "snake tooth") is an extinct genus of synapsid belonging to the family Ophiacodontidae that lived from the Late Carboniferous to the Early Permian in North America and Europe. The genus was named along with its type species ''O. mirus'' by paleontologist Othniel Charles Marsh in 1878 and currently includes five other species. As an ophiacodontid, ''Ophiacodon'' is one of the most basal synapsids and is close to the evolutionary line leading to mammals. Description ''Ophiacodon'' has a large skull with a deep snout. It has the longest skull of any early synapsid, reaching up to in one specimen. The jaws are lined with many small teeth. It was larger than most other tetrapods (four-limbed vertebrates) of its time, ranging from in length and in weight. Specimens of ''Ophiacodon'' vary greatly in size. These differences in size were once used to distinguish species, but are now recognized as ontogenetic variations related to the ages of individuals. S ...
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Echinerpeton
''Echinerpeton'' is an extinct genus of synapsid, including the single species ''Echinerpeton intermedium'' from the Late Carboniferous of Nova Scotia, Canada. The name means 'spiny lizard' (Greek). Along with its contemporary ''Archaeothyris'', ''Echinerpeton'' is the oldest known synapsid, having lived around 308 million years ago. It is known from six small, fragmentary fossils, which were found in an outcrop of the Morien Group near the town of Florence. The most complete specimen preserves articulated vertebrae with high neural spines, indicating that ''Echinerpeton'' was a sail-backed synapsid like the better known '' Dimetrodon'', '' Sphenacodon'', and ''Edaphosaurus''. However, the relationship of ''Echinerpeton'' to these other forms is unclear, and its phylogenetic placement among basal synapsids remains uncertain. Description ''Echinerpeton'' is known from six specimens, five housed in the Museum of Comparative Zoology and a sixth in the Redpath Museum: the holotype M ...
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Caseasauria
Caseasauria is one of the two main clades of early synapsids, the other being the Eupelycosauria. Caseasaurs are currently known only from the Late Carboniferous and the Permian, and include two superficially different families, the small insectivorous or carnivorous Eothyrididae, and the large, herbivorous Caseidae. These two groups share a number of specialised features associated with the morphology of the snout and external naris. The ancestor of caseasaurs can be traced back to an insect eating or an omnivorous reptile-like synapsid from the Pennsylvanian time of the Carboniferous, possibly resembling '' Archaeothyris'', the earliest known synapsid. The caseasaurs were abundant during the later part of the Early Permian, but by the Middle Permian caseasaur diversity declined because the group was outcompeted by the more successful therapsids. The last caseasaurs became extinct at the end of the Guadelupian (Middle Permian). Description Among the most conspicuous char ...
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Ophiacodon Retroversus
''Ophiacodon'' (meaning "snake tooth") is an extinct genus of synapsid belonging to the family Ophiacodontidae that lived from the Late Carboniferous to the Early Permian in North America and Europe. The genus was named along with its type species ''O. mirus'' by paleontologist Othniel Charles Marsh in 1878 and currently includes five other species. As an ophiacodontid, ''Ophiacodon'' is one of the most basal synapsids and is close to the evolutionary line leading to mammals. Description ''Ophiacodon'' has a large skull with a deep snout. It has the longest skull of any early synapsid, reaching up to in one specimen. The jaws are lined with many small teeth. It was larger than most other tetrapods (four-limbed vertebrates) of its time, ranging from in length and in weight. Specimens of ''Ophiacodon'' vary greatly in size. These differences in size were once used to distinguish species, but are now recognized as ontogenetic variations related to the ages of individuals. S ...
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