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Nektaspida
Nektaspida (also called Naraoiida, Nektaspia and Nectaspida) is an extinct order of non- mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae. Naming history and taxonomic placement The order was originally proposed by Raymond in 1920 as Nektaspia. Størmer corrected it to Nectaspida for the 1959 ''Treatise on Invertebrate Paleontology'' to conform with the names of the other trilobite orders. Whittington described it in 1985 with the spelling Nektaspida; the revised 1997 Treatise by Raymond and Fortey uses this spelling, as do other modern works. Whittington (1985) placed the order in the Trilobita. Cotton & Braddy (2000) place it in a new "Trilobite clade" containing the Trilobita, reco ...
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Nektaspida Guts
Nektaspida (also called Naraoiida, Nektaspia and Nectaspida) is an extinct order of non- mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae. Naming history and taxonomic placement The order was originally proposed by Raymond in 1920 as Nektaspia. Størmer corrected it to Nectaspida for the 1959 ''Treatise on Invertebrate Paleontology'' to conform with the names of the other trilobite orders. Whittington described it in 1985 with the spelling Nektaspida; the revised 1997 Treatise by Raymond and Fortey uses this spelling, as do other modern works. Whittington (1985) placed the order in the Trilobita. Cotton & Braddy (2000) place it in a new "Trilobite clade" containing the Trilobita, reco ...
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Buenaspis
''Buenaspis'' is a genus of small ( long) nektaspid arthropod, that lived during the early Cambrian period.Budd, G.E (1999).A Nectaspid Arthropod from the Early Cambrian Sirius Passet Fauna, with a description of Retrodeformation based on Functional Morphology. ''Palaeontology'', 42(1):99–122 Fossil remains of ''Buenaspis'' were collected from the Lower Cambrian Sirius Passet ''Lagerstätte'' of North Greenland. ''Buenaspis'' looks like a soft eyeless trilobite. It has a headshield (or cephalon) slightly larger than the tailshield (pygidium), and in between them six thoracic body segments (somites). The genus is monotypic, its sole species being ''Buenaspis forteyi''. Etymology The name of the genus is derived from the Buen Formation, the deposit where the species was collected, and the Greek word ''aspis'' (shield). The species was named in honor of Richard Fortey, a famed paleontologist. Description ''Buenaspis forteyi'' is between 1 and 3 cm along the axis, a ...
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Artiopoda
The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Hou and Bergström used the name Lamellipedia as a superclass to replace Trilobitomorpha that was originally erected at the subphylum level, which they considered inappropriate. Trilobites, in part due to their mineralising exoskeletons, are by far the most diverse and long lived members of the clade, with most records of other members, which lack mineralised exoskeletons, being from Cambrian deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods, with the proximal lobe being elongate and bearing a lamella, while the distal lobe is paddle-shaped and setiforous (bearing hair-or bristle like structures). The limb endopod has ...
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Artiopodan
The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Hou and Bergström used the name Lamellipedia as a superclass to replace Trilobitomorpha that was originally erected at the subphylum level, which they considered inappropriate. Trilobites, in part due to their mineralising exoskeletons, are by far the most diverse and long lived members of the clade, with most records of other members, which lack mineralised exoskeletons, being from Cambrian deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods, with the proximal lobe being elongate and bearing a lamella, while the distal lobe is paddle-shaped and setiforous (bearing hair-or bristle like structures). The limb endopod has s ...
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Trilobitomorpha
The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Hou and Bergström used the name Lamellipedia as a superclass to replace Trilobitomorpha that was originally erected at the subphylum level, which they considered inappropriate. Trilobites, in part due to their mineralising exoskeletons, are by far the most diverse and long lived members of the clade, with most records of other members, which lack mineralised exoskeletons, being from Cambrian deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods, with the proximal lobe being elongate and bearing a lamella, while the distal lobe is paddle-shaped and setiforous (bearing hair-or bristle like structures). The limb endopod has s ...
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Naraoia
''Naraoia'' is a genus of small to average size (about 2-4½ cm long) marine arthropods within the family Naraoiidae, that lived from the early Cambrian to the late Silurian period. The species are characterized by a large alimentary system and sideways oriented antennas. Etymology The name is derived from Narao, the name of a group of small lakes in Cataract Brook canyon, above Hector on the Canadian Pacific Railway, British Columbia, Canada.Walcott, C.D. Middle Cambrian Branchiopoda, Malacostraca, Trilobita, and Merostomata. In: Cambrian Geology and Paleontology II. Smithsonian. 1914. s:Cambrian Geology and Paleontology/Volume 2/Middle Cambrian Branchiopoda, Malacostraca, Trilobita, and Merostomata History of the classification When the fossil was first discovered in Canada's Burgess Shale, it was believed to be a crustacean, such was the difference between this and other trilobites. Its continuous shield hid most of its structure, interfering with proper classification. ...
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Emucarididae
Emucarididae is an extinct family of soft-shelled trilobite-like arthropods (nektaspids) from the Lower Cambrian of South Australia and South China. It contains only two genera – '' Emucaris'' and '' Kangacaris''. Two species were described in 2010 from specimens recovered from Emu Bay Shale ''Lagerstätte'', one species in 2012 from the Maotianshan Shales. It is classified under the order Nektaspida, and is a sister-group to the families Liwiidae and Naraoiidae. Description The Emucarididae have a non-calcified exoskeleton that consists of an articulating head shield (or cephalon), thorax and tail shield (or pygidium), and there are no constrictions where these parts meet. The cephalon is semi-circular and has a straight back margin. The thorax consists of 3 or 4 narrow segments. The pygidium is 1-2× as long as the cephalon and has a distinct border furrow. The mouth plate (or hypostome In zoology, the hypostome can refer to structures in distinct animal groups: * Hypost ...
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Liwiidae
Liwiidae is a family of arthropods in the order Nektaspida. Members are known from the Cambrian and Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ... periods. Taxonomy *'' Liwia'' Dzik and Lendzion, 1988 :*'' Liwia plana'' Lendzion, 1975 :*'' Liwia convexa'' Lendzion, 1975 *'' Soomaspis'' Fortey & Theron, 1995 :*'' Soomaspis splendida'' Fortey & Theron, 1995 *'' Tariccoia'' Hammann ''et al.'', 1990 :*'' Tariccoia arrusensis'' Hammann ''et al.'', 1990 References Nektaspida Prehistoric arthropod families Cambrian first appearances Ordovician first appearances {{nektaspida-stub ...
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Naraoiidae
Naraoiidae is a family, of extinct, soft-shelled trilobite-like arthropods, that belongs to the order Nectaspida. Species included in the Naraoiidae are known from the second half of the Lower Cambrian to the end of the Upper Silurian. The total number of collection sites is limited and distributed over a vast period of time: Maotianshan Shale and Balang Formation (China), Burgess Shale and Bertie Formation (Canada), the Šárka Formation (Czech Republic), Emu Bay Shale (Australia), Idaho and Utah (USA). This is probably due to the rare occurrence of the right circumstances for soft tissue preservation, needed for these non-calcified exoskeletons. Ecology Naraoiids probably were deposit feeders (''Naraoia'' and '' Pseudonaraoia''), predators or scavengers (''Misszhouia''), living on the sea floor. Description The species of the family ''Naraoiidae'' are almost flat (dorso-ventrally). The upper (or dorsal) side of the body consists of a non-calcified transversely oval or s ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. T ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. T ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is ...
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