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Lycopsis
''Lycopsis'' is an extinct genus of South American metatherian, that lived during the Miocene in Argentina and Colombia. History Although not named until 1927, Florentino Ameghino described a species now seen as synonymous with ''Lycopsis torresi'', ''Anatherium oxyrhynchus'', in 1895 based on a mandibular ramus with several teeth.Marshall, L. G. (1977). A new species of Lycopsis (Borhyaenidae: Marsupialia) from the La Venta fauna (Late Miocene) of Colombia, South America. ''Journal of Paleontology'', 633-642. The fossil was recovered from Puesto Estancia La Costa in Santa Cruz, Argentina, dating to the Miocene. The type material of ''Lycopsis'' was collected in July 1895 by "C. Berry" from the middle Miocene strata of the Santa Cruz Formation along the Santa Cruz River in the same area.Cabrera, Á. (1927). Datos para el conocimiento de los dasiuroideos fósiles argentinos. ''Revista del Museo de la Plata'', ''30'', 271-315. The fossils (MLP 11-113) were fragmentary, constitu ...
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Lycopsis Longirostris
''Lycopsis'' is an extinct genus of South American metatherian, that lived during the Miocene in Argentina and Colombia. History Although not named until 1927, Florentino Ameghino described a species now seen as synonymous with ''Lycopsis torresi'', ''Anatherium oxyrhynchus'', in 1895 based on a mandibular ramus with several teeth.Marshall, L. G. (1977). A new species of Lycopsis (Borhyaenidae: Marsupialia) from the La Venta fauna (Late Miocene) of Colombia, South America. ''Journal of Paleontology'', 633-642. The fossil was recovered from Puesto Estancia La Costa in Santa Cruz, Argentina, dating to the Miocene. The type material of ''Lycopsis'' was collected in July 1895 by "C. Berry" from the middle Miocene strata of the Santa Cruz Formation along the Santa Cruz River in the same area.Cabrera, Á. (1927). Datos para el conocimiento de los dasiuroideos fósiles argentinos. ''Revista del Museo de la Plata'', ''30'', 271-315. The fossils (MLP 11-113) were fragmentary, constitu ...
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Sparassodonta
Sparassodonta (from Greek to tear, rend; and , gen. , ' tooth) is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during th ...
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Borhyaenoidea
Sparassodonta (from Greek to tear, rend; and , gen. , ' tooth) is an extinct order of carnivorous metatherian mammals native to South America, related to modern marsupials. They were once considered to be true marsupials, but are now thought to be a separate side branch that split before the last common ancestor of all modern marsupials. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the ...
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Chasicoan
The Chasicoan ( es, Chasiquense or es, Chasicoense) age is a period of geologic time from 10–9 Ma within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Mayoan and precedes the Huayquerian age.Chasicoan
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Etymology

The Chasicoan is named after the
Arroyo Chasicó Formation Arroyo often refers to: * Arroyo (creek), an intermittently dry creek Arroyo may also refer to: People * Arroyo (surname) Places United States ...
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Santacrucian
The Santacrucian age is a period of geologic time (17.5 – 16.3 Mya (unit), Ma) within the Early Miocene epoch of the Neogene, used more specifically with South American land mammal age, SALMA classification in South America. It follows the Colhuehuapian and precedes the Friasian age. Etymology The age is named after the Santa Cruz Formation in the Magallanes Basin, Austral/Magallanes Basin of southern Patagonia, Argentina and Chile. Formations Fossils References Bibliography

;Santa Cruz Formation * * * * * * * * * * * * * * * * ;Aisol Formation * * ;Cantaure Formation * ;Castillo Formation * * * * * ;Cerro Boleadores Formation * ;Chaguaramas Formation * ;Chilcatay Formation * * * * ;Cura-Mallín Group * * * ;Gran Bajo del Gualicho Formation * ;Jimol Formation * * * * * * * * ;Mariño Formation * * * * * ;Pinturas Formation * * ;Río Yuca Formation * {{SALMA Santacrucian, Miocene South America ...
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Metatheria
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. There are three extant subclasses of mammals, one being metatherians: #monotremes: egg laying mammals like the platypus and the echidna, #metatheria: marsupials, which includes three American orders ( Didelphimorphia, Paucituberculata and Microbiotheria) and four Australasian orders ( Notoryctemorphia, Dasyuromorphia, Peramelemorphia and Diprotodontia), and the # eutherians: placental mammals, consisting of four superorders divided into 21 orders. Metatherians belong to a subgroup of the northern tribosphenic mammal clade or Boreosphenida. They differ from all other mammals in certain morphologies like their dental formula, which includes about five upper and four lower incisors, a canine, three ...
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Santa Cruz Formation, Argentina
The Santa Cruz Formation is a geological formation in the Magallanes/Austral Basin in southern Patagonia in Argentina and in adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with eponymous Santacrucian SALMA. The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which has a lithology predominantly consisting of tuffaceous deposits and fine grained sedimentary claystone and mudstone, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily claystone, mudstone, and sandstone. The environment of deposition is interpreted to have been mostly fluvial, with the lowermost part of the Estancia La Costa Member being transitional between fluvial and marine conditions. The environment of the Estancia La Costa Member is thought to have been relatively warm and humid, but likely became somewhat coole ...
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Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about an ...
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Hondadelphys
''Hondadelphys'' is an extinct genus of carnivorous sparassodonts, known from the Middle Miocene of Colombia. The type species, ''H. fieldsi'', was described in 1976 from the fossil locality of La Venta, which hosts fossils from the Villavieja Formation. ''Hondadelphys ''was originally interpreted as belonging to the opossum family Didelphidae, but subsequently assigned to its own family, Hondadelphidae and interpreted as a basal sparassodont. The genus name refers to the Honda Group, the stratigraphic group in which the fossils of this animal were first found, combined with ''delphys ''(Greek for "womb", a common suffix used for opossum-like metatherian Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well ...s). References Sparassodonts Miocene mammals of South America Lav ...
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