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Hominini
The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The term was originally introduced by Camille Arambourg (1948). Arambourg combined the categories of ''Hominina'' and ''Simiina'' due to Gray (1825) into his new subtribe. Traditionally, chimpanzees, gorillas and orangutans were grouped together as pongids. Since Gray's classification, evidence has accumulated from genetic phylogeny confirming that humans, chimpanzees, and gorillas are more closely related to each other than to the orangutan. The former pongids were reassigned to the subfamily Hominidae ("great apes"), which already included humans, but the details of this reassignment remain contested; within Hominini, not every source excludes gorillas, and not every source includes chimpanzees. Humans are the only extant species in t ...
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Hominidae
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the eastern and western gorilla); '' Pan'' (the chimpanzee and the bonobo); and ''Homo'', of which only modern humans remain. Several revisions in classifying the great apes have caused the use of the term ''"hominid"'' to vary over time. The original meaning of "hominid" referred only to humans (''Homo'') and their closest extinct relatives. However, by the 1990s humans, apes, and their ancestors were considered to be "hominids". The earlier restrictive meaning has now been largely assumed by the term ''"hominin"'', which comprises all members of the human clade after the split from the chimpanzees (''Pan''). The current meaning of "hominid" includes all the great apes including humans. Usage still varies, however, and some scientists and la ...
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Homininae
Homininae (), also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the tribe Hominini (with the genus ''Homo'' including modern humans and numerous extinct species; the subtribe Hominina, comprising at least two extinct genera; and the subtribe Panina, represented only by the genus ''Pan'', which includes chimpanzees and bonobos)―and 2) the tribe Gorillini (gorillas). Alternatively, the genus '' Pan'' is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans (subfamily Ponginae) split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas (through the tribe Gorillini), and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina (see the evolutionary tree below). There are two living species of Panina (chimpanzees and bonobos) and two livin ...
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Homo
''Homo'' () is the genus that emerged in the (otherwise extinct) genus '' Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely related to modern humans (depending on the species), most notably '' Homo erectus'' and '' Homo neanderthalensis''. The genus emerged with the appearance of '' Homo habilis'' just over 2 million years ago. ''Homo'', together with the genus '' Paranthropus'', is probably sister to '' Australopithecus africanus'', which itself had previously split from the lineage of '' Pan'', the chimpanzees. '' Homo erectus'' appeared about 2 million years ago and, in several early migrations, spread throughout Africa (where it is dubbed '' Homo ergaster'') and Eurasia. It was likely that the first human species lived in a hunter-gatherer society and was able to control fire. An adaptive and successful species, ''Homo erectus'' persisted for more than a mi ...
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Graecopithecus
''Graecopithecus'' is an extinct species of hominid that lived in southeast Europe during the late Miocene around 7.2 million years ago. Originally identified by a single lower jaw bone bearing a molar tooth found in Pyrgos Vasilissis, Athens, Greece, in 1944, other tooth specimens were discovered from Azmaka quarry in Bulgaria in 2012. With only little and badly preserved materials to reveal its nature, it is considered as "the most poorly known European Miocene hominoids." The creature was popularly nicknamed 'El Graeco' (word play on the Spanish painter El Greco) by scientists. In 2017, an international team of palaeontologists led by Madelaine Böhme of the Eberhard-Karls-University Tübingen, Germany, published a detailed analysis of the teeth and age of the specimens, and came to the conclusion that it could be the oldest hominin, meaning that it could be the oldest direct ancestors of humans after splitting from that of the chimpanzees. Their simultaneous study also cla ...
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Hominoid Taxonomy 7
Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister group Cercopithecidae form the catarrhine clade, cladistically making them monkeys (though this is the subject of much debate). Apes do not have tails due to a mutation of the TBXT gene. In traditional and non-scientific use, the term "ape" can include tailless primates taxonomically considered Cercopithecidae (such as the Barbary ape and black ape), and is thus not equivalent to the scientific taxon Hominoidea. There are two extant branches of the superfamily Hominoidea: the gibbons, or lesser apes; and the hominids, or great apes. * The family Hylobatidae, the lesser apes, include four genera and a total of 20 species of gibbon, including the lar gibbon and the siamang, all native to Asia. They are highly arboreal and bipedal on the g ...
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Hominina
Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genera ''Homo'', ''Paranthropus'', and ''Kenyanthropus''), and it typically includes the earlier '' Ardipithecus'', '' Orrorin'', '' Sahelanthropus'', and '' Graecopithecus''. All these closely related species are now sometimes collectively termed australopiths or homininians. They are the extinct, close relatives of humans and, with the extant genus ''Homo'', comprise the human clade. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are now called Hominina (''see Hominidae; terms "hominids" and hominins''). While none of the groups normally directly assigned to this group survived, the australopiths do not appear to be literally extinct (in the sense of having no living descendants) as the genera ''Kenyanthropus'', ''Paranthropus'' and ''Homo'' probably emerged as sister of a late ''Aus ...
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Australopithecina
Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genera ''Homo'', ''Paranthropus'', and ''Kenyanthropus''), and it typically includes the earlier '' Ardipithecus'', '' Orrorin'', '' Sahelanthropus'', and '' Graecopithecus''. All these closely related species are now sometimes collectively termed australopiths or homininians. They are the extinct, close relatives of humans and, with the extant genus ''Homo'', comprise the human clade. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are now called Hominina (''see Hominidae; terms "hominids" and hominins''). While none of the groups normally directly assigned to this group survived, the australopiths do not appear to be literally extinct (in the sense of having no living descendants) as the genera ''Kenyanthropus'', ''Paranthropus'' and ''Homo'' probably emerged as sister of a late ''Aus ...
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Australopithecine
Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' ( cladistically including the genera ''Homo'', '' Paranthropus'', and ''Kenyanthropus''), and it typically includes the earlier '' Ardipithecus'', '' Orrorin'', '' Sahelanthropus'', and '' Graecopithecus''. All these closely related species are now sometimes collectively termed australopiths or homininians. They are the extinct, close relatives of humans and, with the extant genus ''Homo'', comprise the human clade. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are now called Hominina (''see Hominidae; terms "hominids" and hominins''). While none of the groups normally directly assigned to this group survived, the australopiths do not appear to be literally extinct (in the sense of having no living descendants) as the genera ''Kenyanthropus'', ''Paranthropus'' and ''Homo'' probably emerged as sister of a late ...
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Sahelanthropus
''Sahelanthropus tchadensis'' is an extinct species of the Homininae (African apes) dated to about , during the Miocene epoch. The species, and its genus ''Sahelanthropus'', was announced in 2002, based mainly on a partial cranium, nicknamed ''Toumaï'', discovered in northern Chad. ''Sahelanthropus tchadensis'' lived close to the time of the chimpanzee–human divergence, possibly related to '' Orrorin'', a species of Homininae that lived about one million years later. It may have been ancestral to both humans and chimpanzees (which would place it in the tribe Hominini), or alternatively an early member of the tribe Gorillini. In 2020, the femur was analyzed, and it was found that ''Sahelanthropus'' was not bipedal, casting some doubt on its position as a human ancestor, but this was refuted in 2022. Taxonomy Discovery Funded by the Mission Paléoanthropologique Franco–Tchadienne, Ahounta Djimdoumalbaye, Fanoné Gongdibé, Mahamat Adoum, and Alain Beauvilain identified t ...
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Human
Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, and language. Humans are highly social and tend to live in complex social structures composed of many cooperating and competing groups, from families and kinship networks to political states. Social interactions between humans have established a wide variety of values, social norms, and rituals, which bolster human society. Its intelligence and its desire to understand and influence the environment and to explain and manipulate phenomena have motivated humanity's development of science, philosophy, mythology, religion, and other fields of study. Although some scientists equate the term ''humans'' with all members of the genus '' Homo'', in common usage, it generally refers to ''Homo sapiens'', the only extant member. Ana ...
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Tribe (biology)
In biology, a tribe is a taxonomic rank above genus, but below family and subfamily. It is sometimes subdivided into subtribes. By convention, all taxonomic ranks from genus upwards are capitalized, including both tribe and subtribe. In zoology, the standard ending for the name of a zoological tribe is "-ini". Examples include the tribes Caprini (goat-antelopes), Hominini (hominins), Bombini (bumblebees), and Thunnini (tunas). The tribe Hominini is divided into subtribes by some scientists; subtribe Hominina then comprises "humans". The standard ending for the name of a zoological subtribe is "-ina". In botany, the standard ending for the name of a botanical tribe is "-eae". Examples include the tribes Acalypheae and Hyacintheae Hyacintheae is a flowering plant tribe in the subfamily Scilloideae Scilloideae (named after the genus ''Scilla'', "squill") is a subfamily of bulbous plants within the family ''Asparagaceae''. Scilloideae is sometimes treated as a separat ...
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Ardipithecus
''Ardipithecus'' is a genus of an extinct hominine that lived during the Late Miocene and Early Pliocene epochs in the Afar Depression, Ethiopia. Originally described as one of the earliest ancestors of humans after they diverged from the chimpanzees, the relation of this genus to human ancestors and whether it is a hominin is now a matter of debate. Two fossil species are described in the literature: '' A. ramidus'', which lived about 4.4 million years ago during the early Pliocene, and '' A. kadabba'', dated to approximately 5.6 million years ago (late Miocene). Initial behavioral analysis indicated that ''Ardipithecus'' could be very similar to chimpanzees, however more recent analysis based on canine size and lack of canine sexual dimorphism indicates that ''Ardipithecus'' was characterised by reduced aggression, and that they more closely resemble bonobos. Some analyses describe ''Australopithecus'' as being sister to ''Ardipithecus ramidus'' specifically. This ...
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