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Evolutionarily Stable Strategy
An evolutionarily stable strategy (ESS) is a strategy (or set of strategies) that is ''impermeable'' when adopted by a population in adaptation to a specific environment, that is to say it cannot be displaced by an alternative strategy (or set of strategies) which may be novel or initially rare. Introduced by John Maynard Smith and George R. Price in 1972/3, it is an important concept in behavioural ecology, evolutionary psychology, mathematical game theory and economics, with applications in other fields such as anthropology, philosophy and political science. In game-theoretical terms, an ESS is an equilibrium refinement of the Nash equilibrium, being a Nash equilibrium that is also "evolutionarily stable." Thus, once fixed in a population, natural selection alone is sufficient to prevent alternative (mutant) strategies from replacing it (although this does not preclude the possibility that a better strategy, or set of strategies, will emerge in response to selective pressure ...
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Nash Equilibrium
In game theory, the Nash equilibrium, named after the mathematician John Nash, is the most common way to define the solution of a non-cooperative game involving two or more players. In a Nash equilibrium, each player is assumed to know the equilibrium strategies of the other players, and no one has anything to gain by changing only one's own strategy. The principle of Nash equilibrium dates back to the time of Cournot, who in 1838 applied it to competing firms choosing outputs. If each player has chosen a strategy an action plan based on what has happened so far in the game and no one can increase one's own expected payoff by changing one's strategy while the other players keep their's unchanged, then the current set of strategy choices constitutes a Nash equilibrium. If two players Alice and Bob choose strategies A and B, (A, B) is a Nash equilibrium if Alice has no other strategy available that does better than A at maximizing her payoff in response to Bob choosing B, and Bo ...
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Anthropology
Anthropology is the scientific study of humanity, concerned with human behavior, human biology, cultures, societies, and linguistics, in both the present and past, including past human species. Social anthropology studies patterns of behavior, while cultural anthropology studies cultural meaning, including norms and values. A portmanteau term sociocultural anthropology is commonly used today. Linguistic anthropology studies how language influences social life. Biological or physical anthropology studies the biological development of humans. Archaeological anthropology, often termed as 'anthropology of the past', studies human activity through investigation of physical evidence. It is considered a branch of anthropology in North America and Asia, while in Europe archaeology is viewed as a discipline in its own right or grouped under other related disciplines, such as history and palaeontology. Etymology The abstract noun '' anthropology'' is first attested in refe ...
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Fisher's Principle
Fisher's principle is an evolutionary model that explains why the sex ratio of most species that produce offspring through sexual reproduction is approximately 1:1 between males and females. A. W. F. Edwards has remarked that it is "probably the most celebrated argument in evolutionary biology". Fisher's principle was outlined by Ronald Fisher in his 1930 book ''The Genetical Theory of Natural Selection'' (but has been incorrectly attributed as original to Fisher). Fisher couched his argument in terms of parental expenditure, and predicted that parental expenditure on both sexes should be equal. Sex ratios that are 1:1 are hence known as "", and those that are ''not'' 1:1 are "" or "" and occur because they break the assumptions made in Fisher's model.Charnov, E., and Bull, J. (1989).Non-fisherian sex ratios with sex change and environmental sex determination, ''Nature 338,'' pp. 148–150. https://doi.org/10.1038/338148a0 Retrieved 29 March 2022. Basic explanation W.D. Ham ...
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Sex Ratio
The sex ratio (or gender ratio) is usually defined as the ratio of males to females in a population. As explained by Fisher's principle, for evolutionary reasons this is typically about 1:1 in species which reproduce sexually. Many species deviate from an even sex ratio, either periodically or permanently. Examples include parthenogenic species, periodically mating organisms such as aphids, some eusocial wasps, bees, ants, and termites. The human sex ratio is of particular interest to anthropologists and demographers. In human societies, sex ratios at birth may be considerably skewed by factors such as the age of mother at birth and by sex-selective abortion and infanticide. Exposure to pesticides and other environmental contaminants may be a significant contributing factor as well. As of 2014, the global sex ratio at birth is estimated at 107 boys to 100 girls (1,000 boys per 934 girls).. Types In most species, the sex ratio varies according to the age profile of the ...
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Robert MacArthur
Robert Helmer MacArthur (April 7, 1930 – November 1, 1972) was a Canadian-born American ecologist who made a major impact on many areas of community and population ecology. Early life and education MacArthur was born in Toronto, Ontario, to John Wood MacArthur and Olive Turner in 1930. He later moved to Marlboro, Vermont, as his father moved from the University of Toronto to Marlboro College. MacArthur received his Bachelor's degree in mathematics from Marlboro College, followed by a Master's degree in mathematics from Brown University in 1953. A student of G. Evelyn Hutchinson, MacArthur earned his Ph.D. from Yale University in 1957; his thesis was on the division of ecological niches among five warbler species in the conifer forests of Maine and Vermont. From 1957 to 1958, MacArthur worked as a postdoc with David Lack. Career MacArthur was a professor at the University of Pennsylvania, 1958–65, and professor of biology at Princeton University, 1965-72. He played a ...
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Evolution And The Theory Of Games
''Evolution and the Theory of Games'' is a book by the British evolutionary biologist John Maynard Smith on evolutionary game theory. The book was initially published in December 1982 by Cambridge University Press. Overview In the book, John Maynard Smith summarises work on evolutionary game theory that had developed in the 1970s, to which he made several important contributions. The book is also noted for being well written and not overly mathematically challenging. The main contribution to be had from this book is the introduction of the Evolutionarily Stable Strategy, or ESS, concept, which states that for a set of behaviours to be conserved over evolutionary time, they must be the most profitable avenue of action when common, so that no alternative behaviour can invade. So, for instance, suppose that in a population of frogs, males fight to the death over breeding ponds. This would be an ESS if any one cowardly frog that does not fight to the death always fares worse (in ...
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Journal Of Theoretical Biology
The ''Journal of Theoretical Biology'' is a biweekly peer-reviewed scientific journal covering theoretical biology, as well as mathematical, computational, and statistical aspects of biology. Some research areas covered by the journal include cell biology, evolutionary biology, population genetics, morphogenesis, and immunology. The journal was established in 1961. Its founding editor-in-chief was English biologist James F. Danielli, who remained editor until his death in 1984. The journal is published by Elsevier and, , the editors-in-chief are Denise Kirschner (University of Michigan Medical School), Mark Chaplain ( University of St. Andrews), and Akira Sawaki (Aichi Cancer Center, Nagoya). Lewis Wolpert served as editor-in-chief for more than 55 years. According to the ''Journal Citation Reports'' the journal has a 2020 impact factor of 2.691. Notable articles The following are the most highly cited articles (more than 2000 citations at April 2021) that have been published ...
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Nature (journal)
''Nature'' is a British weekly scientific journal founded and based in London, England. As a multidisciplinary publication, ''Nature'' features peer-reviewed research from a variety of academic disciplines, mainly in science and technology. It has core editorial offices across the United States, continental Europe, and Asia under the international scientific publishing company Springer Nature. ''Nature'' was one of the world's most cited scientific journals by the Science Edition of the 2019 '' Journal Citation Reports'' (with an ascribed impact factor of 42.778), making it one of the world's most-read and most prestigious academic journals. , it claimed an online readership of about three million unique readers per month. Founded in autumn 1869, ''Nature'' was first circulated by Norman Lockyer and Alexander Macmillan as a public forum for scientific innovations. The mid-20th century facilitated an editorial expansion for the journal; ''Nature'' redoubled its efforts in ...
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Mutant
In biology, and especially in genetics, a mutant is an organism or a new genetic character arising or resulting from an instance of mutation, which is generally an alteration of the DNA sequence of the genome or chromosome of an organism. It is a characteristic that would not be observed naturally in a specimen. The term mutant is also applied to a virus with an alteration in its nucleotide sequence whose genome is in the nuclear genome. The natural occurrence of genetic mutations is integral to the process of evolution. The study of mutants is an integral part of biology; by understanding the effect that a mutation in a gene has, it is possible to establish the normal function of that gene. Mutants arise by mutation Mutants arise by mutations occurring in pre-existing genomes as a result of errors of DNA replication or errors of DNA repair. Errors of replication often involve translesion synthesis by a DNA polymerase when it encounters and bypasses a damaged base in the ...
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Natural Selection
Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Charles Darwin popularised the term "natural selection", contrasting it with artificial selection, which in his view is intentional, whereas natural selection is not. Variation exists within all populations of organisms. This occurs partly because random mutations arise in the genome of an individual organism, and their offspring can inherit such mutations. Throughout the lives of the individuals, their genomes interact with their environments to cause variations in traits. The environment of a genome includes the molecular biology in the cell, other cells, other individuals, populations, species, as well as the abiotic environment. Because individuals with certain variants of the trait tend to survive and reproduce more than individual ...
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Fixation (population Genetics)
In population genetics, fixation is the change in a gene pool from a situation where there exists at least two variants of a particular gene (allele) in a given population to a situation where only one of the alleles remains. In the absence of mutation or heterozygote advantage, any allele must eventually be lost completely from the population or fixed (permanently established at 100% frequency in the population). Whether a gene will ultimately be lost or fixed is dependent on selection coefficients and chance fluctuations in allelic proportions. Fixation can refer to a gene in general or particular nucleotide position in the DNA chain ( locus). In the process of substitution, a previously non-existent allele arises by mutation and undergoes fixation by spreading through the population by random genetic drift or positive selection. Once the frequency of the allele is at 100%, i.e. being the only gene variant present in any member, it is said to be "fixed" in the population. Sim ...
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Ecological Stability
In ecology, an ecosystem is said to possess ecological stability (or equilibrium) if it is capable of returning to its equilibrium state after a perturbation (a capacity known as resilience) or does not experience unexpected large changes in its characteristics across time. Although the terms community stability and ecological stability are sometimes used interchangeably, community stability refers only to the characteristics of communities. It is possible for an ecosystem or a community to be stable in some of their properties and unstable in others. For example, a vegetation community in response to a drought might conserve biomass but lose biodiversity. Stable ecological systems abound in nature, and the scientific literature has documented them to a great extent. Scientific studies mainly describe grassland plant communities and microbial communities. Nevertheless, it is important to mention that not every community or ecosystem in nature is stable (for example, wolves and m ...
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