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Elasmobranchii
Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including sharks (superorder Selachii), rays, skates, and sawfish (superorder Batoidea). Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of the clade is unclear with respect to fossil chondrichthyans. It has been used by different authors as equivalent to Neoselachii (the clade including modern sharks and rays and their last common ancestor) or f ...
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Chondrichthyes
Chondrichthyes (; ) is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fishes'', which have skeletons primarily composed of bone tissue. Chondrichthyes are jawed vertebrates with paired fins, paired nares, scales, and a heart with its chambers in series. Extant chondrichthyes range in size from the 10 cm (3.9 in) finless sleeper ray to the 10 m (32 ft) whale shark. The class is divided into two subclasses: Elasmobranchii ( sharks, rays, skates, and sawfish) and Holocephali (chimaeras, sometimes called ghost sharks, which are sometimes separated into their own class). Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates. Anatomy Skeleton The skeleton is cartilaginous. The notochord is gradually replaced by a vertebral column during development, except in Holocephali, where the notochord stays intact. In some d ...
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Shark
Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorpha (or Selachii) and are the sister group to the rays. However, the term "shark" has also been used to refer to all extinct members of Chondrichthyes with a shark-like morphology, such as hybodonts and xenacanths. The oldest modern sharks are known from the Early Jurassic. They range in size from the small dwarf lanternshark (''Etmopterus perryi''), a deep sea species that is only in length, to the whale shark (''Rhincodon typus''), the largest fish in the world, which reaches approximately in length. Sharks are found in all seas and are common to depths up to . They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river shark, which can be found in both seawater ...
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Sawfish
Sawfish, also known as carpenter sharks, are a family of rays characterized by a long, narrow, flattened rostrum, or nose extension, lined with sharp transverse teeth, arranged in a way that resembles a saw. They are among the largest fish with some species reaching lengths of about . They are found worldwide in tropical and subtropical regions in coastal marine and brackish estuarine waters, as well as freshwater rivers and lakes. They are endangered. They should not be confused with sawsharks (order Pristiophoriformes) or the extinct sclerorhynchoids (order Rajiformes) which have a similar appearance, or swordfish (family Xiphiidae) which have a similar name but a very different appearance. Sawfishes are relatively slow breeders and the females give birth to live young. They feed on fish and invertebrates that are detected and captured with the use of their saw. They are generally harmless to humans, but can inflict serious injuries with the saw when captured and defendi ...
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Batoidea
Batoidea is a superorder of cartilaginous fishes, commonly known as rays. They and their close relatives, the sharks, comprise the subclass Elasmobranchii. Rays are the largest group of cartilaginous fishes, with well over 600 species in 26 families. Rays are distinguished by their flattened bodies, enlarged pectoral fins that are fused to the head, and gill slits that are placed on their ventral surfaces. Anatomy Batoids are flat-bodied, and, like sharks, are cartilaginous fish, meaning they have a boneless skeleton made of a tough, elastic cartilage. Most batoids have five ventral slot-like body openings called gill slits that lead from the gills, but the Hexatrygonidae have six. Batoid gill slits lie under the pectoral fins on the underside, whereas a shark's are on the sides of the head. Most batoids have a flat, disk-like body, with the exception of the guitarfishes and sawfishes, while most sharks have a spindle-shaped body. Many species of batoid have developed their p ...
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Holocephali
Holocephali ("complete heads"), sometimes given the term Euchondrocephali, is a subclass of cartilaginous fish in the class Chondrichthyes. The earliest fossils are of teeth and come from the Devonian period. Little is known about these primitive forms, and the only surviving group in the subclass is the order Chimaeriformes. Chimaeriformes, commonly known as chimaeras, includes the rat fishes in the genus ''Chimaera'', and the elephant fishes in the genus ''Callorhinchus''. These fishes move by using sweeping movements of their large pectoral fins. They are deep sea fish with slender tails, living close to the seabed to feed on benthic invertebrates. They lack a stomach, their food moving directly into the intestine. Extinct holocephalans were much more diverse in lifestyles, including shark-like predatory forms and slow, durophagous fish. Characteristics Members of this taxon preserve today some features of elasmobranch life in Paleozoic times, though in other respects they ...
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Hybodontiformes
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like chondrichthyans, which existed from the late Devonian to the Late Cretaceous. They form the group of Elasmobranchii closest to neoselachians, the clade of modern sharks and rays. Hybodonts were named and are distinguished based on their conical tooth shape. They are also noted for the presence of a spine on each of their two dorsal fins. They were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct. Etymology The term hybodont comes from the Greek word ''ὕβος'' or ''ὑβός'' meaning hump or hump-backed and ''ὀδούς, ὀδοντ'' meaning tooth. This name was given based on their conical compr ...
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Great White Shark
The great white shark (''Carcharodon carcharias''), also known as the white shark, white pointer, or simply great white, is a species of large mackerel shark which can be found in the coastal surface waters of all the major oceans. It is notable for its size, with larger female individuals growing to in length and in weight at maturity. However, most are smaller; males measure , and females measure on average. According to a 2014 study, the lifespan of great white sharks is estimated to be as long as 70 years or more, well above previous estimates, making it one of the longest lived cartilaginous fishes currently known. According to the same study, male great white sharks take 26 years to reach sexual maturity, while the females take 33 years to be ready to produce offspring. Great white sharks can swim at speeds of 25 km/h (16 mph) for short bursts and to depths of . The great white shark is an apex predator, as it has no known natural predators other than, on ve ...
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Skate (fish)
Skates are cartilaginous fish belonging to the family Rajidae in the superorder Batoidea of rays. More than 150 species have been described, in 17 genera.LAST, P.R. & SÉRET, B. & STEHMANN, M.F.W. & WEIGMANN, S. (2016) Skates, Family Rajidae. In: Last, P.R., White, W.T., Carvalho, M.R. de, Séret, B., Stehmann, M.F.W & Naylor, G.J.P (Eds.) Rays of the World. CSIRO Publishing, Melbourne: 204–363 Softnose skates and pygmy skates were previously treated as subfamilies of Rajidae (Arhynchobatinae and Gurgesiellinae), but are now considered as distinct families. Alternatively, the name "skate" is used to refer to the entire order of Rajiformes (families Anacanthobatidae, Arhynchobatidae, Gurgesiellidae and Rajidae). Members of Rajidae are distinguished by a stiff snout and a rostrum that is not reduced. Taxonomy and systematics Evolution Skates belong to the ancient lineage of cartilaginous fishes. Fossil denticles (tooth-like scales in the skin) resembling those of today' ...
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Class (biology)
In biological classification, class ( la, classis) is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, and species, with class fitting between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name (and not just called a ''top-level genus'' ''(genus summum)'') was first introduced by the French botanist Joseph Pitton de Tournefort in his classification of plants that appeared in his ''Eléments de botanique'', 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominat ...
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Chimaera
Chimaeras are cartilaginous fish in the order Chimaeriformes , known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively. At one time a "diverse and abundant" group (based on the fossil record), their closest living relatives are sharks and rays, though their last common ancestor with them lived nearly 400 million years ago. Today, they are largely confined to deep water. Description and habits Chimaeras live in temperate ocean floors down to deep, with few occurring at depths shallower than . Exceptions include the members of the genus ''Callorhinchus'', the rabbit fish and the spotted ratfish, which locally or periodically can be found at shallower depths. Consequently, these are also among the few species from the chimaera order kept in public aquaria. They live in all the oceans except for the Arctic and Antarctic oceans. They have elongated, soft ...
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Clasper
In biology, a clasper is a male anatomical structure found in some groups of animals, used in mating. Male cartilaginous fish have claspers formed from the posterior portion of their pelvic fin which serve to channel semen into the female's cloaca during mating. The act of mating in some fish including sharks usually includes one of the claspers raised to allow water into the siphon through a specific orifice. The clasper is then inserted into the cloaca, where it opens like an umbrella to anchor its position. The siphon then begins to contract, expelling water and sperm. The claspers of many shark species have spines or hooks, which may hold them in place during copulation. Male chimaeras have cephalic claspers (tenacula) on their heads, which are thought to aid in holding the female during mating. In entomology, it is a structure in male insects that is used to hold the female during copulation (see Lepidoptera genitalia for more). See also *Sexual coercion among animal ...
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