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Beak
The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, courtship, and feeding young. The terms ''beak'' and ''rostrum'' are also used to refer to a similar mouth part in some ornithischians, pterosaurs, cetaceans, dicynodonts, anuran tadpoles, monotremes (i.e. echidnas and platypuses, which have a beak-like structure), sirens, pufferfish, billfishes and cephalopods. Although beaks vary significantly in size, shape, color and texture, they share a similar underlying structure. Two bony projections – the upper and lower mandibles – are covered with a thin keratinized layer of epidermis known as the rhamphotheca. In most species, two holes called ''nares'' lead to the respiratory system. Etymology Although the word "beak" was, in the past, generally restricted to the sharpened b ...
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Cephalopod Beak
All extant cephalopods have a two-part beak, or rostrum, situated in the buccal mass and surrounded by the muscular head appendages. The dorsal (upper) mandible fits into the ventral (lower) mandible and together they function in a scissor-like fashion.Young, R.E., M. Vecchione & K.M. Mangold (1999)Cephalopoda Glossary Tree of Life Web Project. The beak may also be referred to as the mandibles or jaws.Tanabe, K., Y. Hikida & Y. Iba (2006). Two coleoid jaws from the Upper Cretaceous of Hokkaido, Japan. ''Journal of Paleontology'' 80(1): 138–145. Fossilised remains of beaks are known from a number of cephalopod groups, both extant and extinct, including squids, octopuses, belemnites, and vampyromorphs. Aptychi – paired plate-like structures found in ammonites – may also have been jaw elements. Composition Composed primarily of chitin and cross-linked proteins, beaks are more-or-less indigestible and are often the only identifiable cephalopod remains found in the stomac ...
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Echidna
Echidnas (), sometimes known as spiny anteaters, are quill-covered monotremes (egg-laying mammals) belonging to the family Tachyglossidae . The four extant species of echidnas and the platypus are the only living mammals that lay eggs and the only surviving members of the order Monotremata. The diet of some species consists of ants and termites, but they are not closely related to the true anteaters of the Americas, which (along with sloths and armadillos) are xenarthrans. Echidnas live in Australia and New Guinea. Echidnas evolved between 20 and 50 million years ago, descending from a platypus-like monotreme. This ancestor was aquatic, but echidnas adapted to life on land. Etymology Echidnas are named after Echidna, a creature from Greek mythology who was half-woman, half-snake, as the animal was perceived to have qualities of both mammals and reptiles. An alternative explanation is a confusion with . Physical characteristics Echidnas are medium-sized, solitary ma ...
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Monotreme
Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals ( Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their brains, jaws, digestive tract, reproductive tract, and other body parts, compared to the more common mammalian types. In addition, they lay eggs rather than bearing live young, but, like all mammals, the female monotremes nurse their young with milk. Monotremes have been considered members of Australosphenida, a clade that contains extinct mammals from the Jurassic and Cretaceous of Madagascar, South America, and Australia, though this is disputed. The only surviving examples of monotremes are all indigenous to Australia and New Guinea, although there is evidence that they were once more widespread, as '' Monotrematum'' is known from the Paleocene of South America. The extant monotreme species are the platypus and four species of echidnas. ...
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Cetacea
Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver. While the majority of cetaceans live in marine environments, a small number exclusively reside in brackish water or fresh water. Having a cosmopolitan distribution, they can be found in some rivers and all of Earth's oceans, and many species inhabit vast ranges where they migrate with the changing of the seasons. Cetaceans are famous for their high intelligence and complex social behaviour as well as for the enormous size of some of the group's members, such as the blue whale which reaches a maximum confirmed length of 29.9 meters (98 feet) and a weight of 173 tonnes (190 short t ...
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Trabecula
A trabecula (plural trabeculae, from Latin for "small beam") is a small, often microscopic, tissue element in the form of a small beam, strut or rod that supports or anchors a framework of parts within a body or organ. A trabecula generally has a mechanical function, and is usually composed of dense collagenous tissue (such as the trabecula of the spleen). It can be composed of other material such as muscle and bone. In the heart, muscles form trabeculae carneae and septomarginal trabeculae. Cancellous bone is formed from groupings of trabeculated bone tissue. In cross section, trabeculae of a cancellous bone can look like septa, but in three dimensions they are topologically distinct, with trabeculae being roughly rod or pillar-shaped and septa being sheet-like. When crossing fluid-filled spaces, trabeculae may offer the function of resisting tension (as in the penis, see for example trabeculae of corpora cavernosa and trabeculae of corpus spongiosum) or providing a c ...
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Birds
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the bee hummingbird to the ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimmin ...
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Preening
Preening is a found in birds that involves the use of the beak to position feathers, interlock feather that have become separated, clean plumage, and keep ectoparasites in check. Feathers contribute significantly to a bird's insulation, waterproofing and aerodynamic flight, and so are vital to its survival. Because of this, birds spend considerable time each day maintaining their feathers, primarily through preening. Several actions make up preening behaviour. Birds fluff up and shake their feathers, which helps to "rezip" feather barbules that have become unhooked. Using their beaks, they gather preen oil from a gland at the base of their tail and distribute this oil through their feathers. They draw each contour feather through their bill, nibbling it from base to tip. Over time, some elements of preening have evolved to have secondary functions. Ritualised preening has become a part of some courtship displays, for example. It is also a displacement activity that can occur ...
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Rostrum (anatomy)
Rostrum (from Latin ', meaning ''beak'') is a term used in anatomy for a number of phylogenetically unrelated structures in different groups of animals. Invertebrates * In crustaceans, the rostrum is the forward extension of the carapace in front of the eyes. It is generally a rigid structure, but can be connected by a hinged joint, as seen in Leptostraca. * Among insects, the rostrum is the name for the piercing mouthparts of the order Hemiptera as well as those of the snow scorpionflies, among many others. The long snout of weevils is also called a rostrum. * Gastropod molluscs have a rostrum or proboscis. * Cephalopod molluscs have hard beak-like mouthparts referred to as the rostrum. File:Washington DC Zoo - Macrobrachium rosenbergii 6.jpg, Crustacean: the rostrum of the shrimp '' Macrobrachium rosenbergii'' is serrated along both edges. File:Gminatus australis with Beetle.jpg, Insect: assassin bug piercing its prey with its rostrum File:Architeuthis beak.jpg, Cephal ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were i ...
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Platypus
The platypus (''Ornithorhynchus anatinus''), sometimes referred to as the duck-billed platypus, is a semiaquatic, egg-laying mammal endemic to eastern Australia, including Tasmania. The platypus is the sole living representative or monotypic taxon of its family ( Ornithorhynchidae) and genus (''Ornithorhynchus''), though a number of related species appear in the fossil record. Together with the four species of echidna, it is one of the five extant species of monotremes, mammals that lay eggs instead of giving birth to live young. Like other monotremes, it senses prey through electrolocation. It is one of the few species of venomous mammals, as the male platypus has a spur on the hind foot that delivers a venom, capable of causing severe pain to humans. The unusual appearance of this egg-laying, duck-billed, beaver-tailed, otter-footed mammal baffled European naturalists when they first encountered it, and the first scientists to examine a preserved platypus ...
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Billfish
The term billfish refers to a group of saltwater predatory fish characterised by prominent pointed bills (rostra), and by their large size; some are longer than . Extant billfish include sailfish and marlin, which make up the family Istiophoridae; and swordfish, sole member of the family Xiphiidae. They are apex predators which feed on a wide variety of smaller fish, crustaceans and cephalopods. These two families are sometimes classified as belonging to the order Istiophoriformes, a group which originated around 71 million years ago in the Late Cretaceous, with the two families diverging around 15 million years ago in the Late Miocene. However, they are also classified as being closely related to the mackerels and tuna within the suborder Scombroidei of the order Perciformes. However, the 5th edition of the ''Fishes of the World'' does recognise the Istiophoriformes as a valid order, albeit including the Sphyraenidae, the barracudas. Billfish are pelagic and highly mi ...
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Dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the sk ...
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