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Australopithecines
Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' ( cladistically including the genera ''Homo'', '' Paranthropus'', and ''Kenyanthropus''), and it typically includes the earlier '' Ardipithecus'', '' Orrorin'', '' Sahelanthropus'', and '' Graecopithecus''. All these closely related species are now sometimes collectively termed australopiths or homininians. They are the extinct, close relatives of humans and, with the extant genus ''Homo'', comprise the human clade. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are now called Hominina (''see Hominidae; terms "hominids" and hominins''). While none of the groups normally directly assigned to this group survived, the australopiths do not appear to be literally extinct (in the sense of having no living descendants) as the genera ''Kenyanthropus'', ''Paranthropus'' and ''Homo'' probably emerged as sister of a late ''A ...
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Paranthropus Robustus
''Paranthropus robustus'' is a species of robust australopithecine from the Early and possibly Middle Pleistocene of the Cradle of Humankind, South Africa, about 2.27 to 0.87 (or, more conservatively, 2 to 1) million years ago. It has been identified in Kromdraai, Swartkrans, Sterkfontein, Gondolin, Cooper's, and Drimolen Caves. Discovered in 1938, it was among the first early hominins described, and became the type species for the genus '' Paranthropus''. However, it has been argued by some that ''Paranthropus'' is an invalid grouping and synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus robustus''. Robust australopithecines—as opposed to gracile australopithecines—are characterised by heavily built skulls capable of producing high stresses and bite forces, as well as inflated cheek teeth ( molars and premolars). Males had more heavily built skulls than females. ''P. robustus'' may have had a genetic susceptibility ...
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Australopithecus Sediba
''Australopithecus sediba'' is an extinct species of australopithecine recovered from Malapa Cave, Cradle of Humankind, South Africa. It is known from a partial juvenile skeleton, the holotype MH1, and a partial adult female skeleton, the paratype MH2. They date to about 1.98 million years ago in the Early Pleistocene, and coexisted with ''Paranthropus robustus'' and '' Homo ergaster'' / ''Homo erectus''. Malapa is interpreted as having been a natural death trap, the base of a long vertical shaft which creatures could accidentally fall into. ''A. sediba'' was initially described as being a potential human ancestor, and perhaps the progenitor of ''Homo'', but this is contested and it could also represent a late-surviving population or sister species of '' A. africanus'' which had earlier inhabited the area. MH1 has a brain volume of about 420–440 cc, similar to other australopithecines. The face of MH1 is strikingly similar to ''Homo'' instead of other australopithecines, wit ...
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Paranthropus Boisei
''Paranthropus boisei'' is a species of australopithecine from the Early Pleistocene of East Africa about 2.5 to 1.15 million years ago. The holotype specimen, OH 5, was discovered by palaeoanthropologist Mary Leakey in 1959, and described by her husband Louis a month later. It was originally placed into its own genus as "''Zinjanthropus boisei''", but is now relegated to '' Paranthropus'' along with other robust australopithecines. However, it is also argued that ''Paranthropus'' is an invalid grouping and synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus boisei''. Robust australopithecines are characterised by heavily built skulls capable of producing high stresses and bite forces, and some of the largest molars with the thickest enamel of any known ape. ''P. boisei'' is the most robust of this group. Brain size was about , similar to other australopithecines. Some skulls are markedly smaller than others, which is taken ...
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Australopithecus
''Australopithecus'' (, ; ) is a genus of early hominins that existed in Africa during the Late Pliocene and Early Pleistocene. The genus ''Homo'' (which includes modern humans) emerged within ''Australopithecus'', as sister to e.g. ''Australopithecus sediba''. Also the genera '' Paranthropus'' and ''Kenyanthropus'' emerged within ''Australopithecus''. ''Australopithecus'' is a member of the subtribe Australopithecina, which sometimes also includes '' Ardipithecus'', though the term "australopithecine" is sometimes used to refer only to members of ''Australopithecus''. Species include '' A. garhi'', '' A. africanus'', '' A. sediba'', ''A. afarensis, A. anamensis, A. bahrelghazali'' and '' A. deyiremeda''. Debate exists as to whether some ''Australopithecus'' species should be reclassified into new genera, or if ''Paranthropus'' and ''Kenyanthropus'' are synonymous with ''Australopithecus'', in part because of the taxonomic inconsistency. The earliest known member of the genu ...
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Paranthropus
''Paranthropus'' is a genus of extinct hominin which contains two widely accepted species: '' P. robustus'' and '' P. boisei''. However, the validity of ''Paranthropus'' is contested, and it is sometimes considered to be synonymous with ''Australopithecus''. They are also referred to as the robust australopithecines. They lived between approximately 2.6 and 1.2 million years ago (mya) from the end of the Pliocene to the Middle Pleistocene. ''Paranthropus'' is characterised by robust skulls, with a prominent gorilla-like sagittal crest along the midline—which suggest strong chewing muscles—and broad, herbivorous teeth used for grinding. However, they likely preferred soft food over tough and hard food. ''Paranthropus'' species were generalist feeders, but ''P. robustus'' was likely an omnivore, whereas ''P. boisei'' was likely herbivorous and mainly ate bulbotubers. They were bipeds. Despite their robust heads, they had comparatively small bodies. Average weight and heig ...
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Praeanthropus
''Australopithecus'' (, ; ) is a genus of early hominins that existed in Africa during the Late Pliocene and Early Pleistocene. The genus ''Homo'' (which includes modern humans) emerged within ''Australopithecus'', as sister to e.g. ''Australopithecus sediba''. Also the genera ''Paranthropus'' and ''Kenyanthropus'' emerged within ''Australopithecus''. ''Australopithecus'' is a member of the subtribe Australopithecina, which sometimes also includes '' Ardipithecus'', though the term "australopithecine" is sometimes used to refer only to members of ''Australopithecus''. Species include '' A. garhi'', '' A. africanus'', '' A. sediba'', ''A. afarensis, A. anamensis, A. bahrelghazali'' and '' A. deyiremeda''. Debate exists as to whether some ''Australopithecus'' species should be reclassified into new genera, or if ''Paranthropus'' and ''Kenyanthropus'' are synonymous with ''Australopithecus'', in part because of the taxonomic inconsistency. The earliest known member of the genus, ...
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Australopithecus Afarensis
''Australopithecus afarensis'' is an extinct species of australopithecine which lived from about 3.9–2.9 million years ago (mya) in the Pliocene of East Africa. The first fossils were discovered in the 1930s, but major fossil finds would not take place until the 1970s. From 1972 to 1977, the International Afar Research Expedition—led by anthropologists Maurice Taieb, Donald Johanson and Yves Coppens—unearthed several hundreds of hominin specimens in Hadar, Ethiopia, the most significant being the exceedingly well-preserved skeleton AL 288-1 (" Lucy") and the site AL 333 ("the First Family"). Beginning in 1974, Mary Leakey led an expedition into Laetoli, Tanzania, and notably recovered fossil trackways. In 1978, the species was first described, but this was followed by arguments for splitting the wealth of specimens into different species given the wide range of variation which had been attributed to sexual dimorphism (normal differences between males and females) ...
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Australopithecus Africanus
''Australopithecus africanus'' is an extinct species of australopithecine which lived between about 3.3 and 2.1 million years ago in the Late Pliocene to Early Pleistocene of South Africa. The species has been recovered from Taung, Sterkfontein, Makapansgat, and Gladysvale. The first specimen, the Taung child, was described by anatomist Raymond Dart in 1924, and was the first early hominin found. However, its closer relations to humans than to other apes would not become widely accepted until the middle of the century because most had believed humans evolved outside of Africa. It is unclear how ''A. africanus'' relates to other hominins, being variously placed as ancestral to ''Homo'' and '' Paranthropus'', to just ''Paranthropus'', or to just '' P. robustus''. The specimen " Little Foot" is the most completely preserved early hominin, with 90% of the skeleton intact, and the oldest South African australopith. However, it is controversially suggested that it and similar spe ...
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Kenyanthropus
''Kenyanthropus'' is a hominin genus identified from the Lomekwi site by Lake Turkana, Kenya, dated to 3.3 to 3.2 million years ago during the Middle Pliocene. It contains one species, ''K. platyops'', but may also include the 2 million year old '' Homo rudolfensis'', or ''K. rudolfensis''. Before its naming in 2001, '' Australopithecus afarensis'' was widely regarded as the only australopithecine to exist during the Middle Pliocene, but ''Kenyanthropus'' evinces a greater diversity than once acknowledged. ''Kenyanthropus'' is most recognisable by an unusually flat face and small teeth for such an early hominin, with values on the extremes or beyond the range of variation for australopithecines in regard to these features. Multiple australopithecine species may have coexisted by foraging for different food items ( niche partitioning), which may be reason why these apes anatomically differ in features related to chewing. The Lomekwi site also yielded the earliest stone tool ...
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Hominini
The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The term was originally introduced by Camille Arambourg (1948). Arambourg combined the categories of ''Hominina'' and ''Simiina'' due to Gray (1825) into his new subtribe. Traditionally, chimpanzees, gorillas and orangutans were grouped together as pongids. Since Gray's classification, evidence has accumulated from genetic phylogeny confirming that humans, chimpanzees, and gorillas are more closely related to each other than to the orangutan. The former pongids were reassigned to the subfamily Hominidae ("great apes"), which already included humans, but the details of this reassignment remain contested; within Hominini, not every source excludes gorillas, and not every source includes chimpanzees. Humans are the only extant species in t ...
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Australopithecus Garhi
''Australopithecus garhi'' is a species of australopithecine from the Bouri Formation in the Afar Region of Ethiopia 2.6–2.5 million years ago (mya) during the Early Pleistocene. The first remains were described in 1999 based on several skeletal elements uncovered in the three years preceding. ''A. garhi'' was originally considered to have been a direct ancestor to ''Homo'' and the human line, but is now thought to have been an offshoot. Like other australopithecines, ''A. garhi'' had a brain volume of ; a jaw which jutted out (prognathism); relatively large molars and premolars; adaptations for both walking on two legs (bipedalism) and grasping while climbing (arboreality); and it is possible that, though unclear if, males were larger than females (exhibited sexual dimorphism). One individual, presumed female based on size, may have been tall. ''A. garhi'' is the first pre-''Homo'' hominin postulated to have manufactured tools—using them in butchering—and may be counted ...
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Paranthropus Aethiopicus
''Paranthropus aethiopicus'' is an extinct species of robust australopithecine from the Late Pliocene to Early Pleistocene of East Africa about 2.7–2.3 million years ago. However, it is much debated whether or not ''Paranthropus'' is an invalid grouping and is synonymous with ''Australopithecus'', so the species is also often classified as ''Australopithecus aethiopicus''. Whatever the case, it is considered to have been the ancestor of the much more robust '' P. boisei''. It is debated if ''P. aethiopicus'' should be subsumed under ''P. boisei'', and the terms ''P. boisei'' sensu lato ("in the broad sense") and ''P. boisei'' sensu stricto ("in the strict sense") can be used to respectively include and exclude ''P. aethiopicus'' from ''P. boisei''. Like other ''Paranthropus'', ''P. aethiopicus'' had a tall face, thick palate, and especially enlarged cheek teeth. However, likely due to its archaicness, it also diverges from other ''Paranthropus'', with some aspects resembling th ...
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