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Archaeoceti
Archaeoceti ("ancient whales"), or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene (). Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia , resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation . All archaeocetes from the Ypresian (56–47.8 mya) and most from the Lutetian (47.8–41.3 mya) are known exclusively from Indo-Pakistan, but Bartonian (41.3–38.0 mya) and Priabonian (38.0–33.9 mya) genera are known from across Earth, including North America, Egypt, New Zealand, and Europe. Although no consensus exists regarding the mode of locomotion of which cetaceans were capable during the ...
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Ambulocetus Fossil Remains
''Ambulocetus'' ( Latin ''ambulare'' "to walk" + ''cetus'' "whale") is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene ( Lutetian). It contains one species, ''Ambulocetus natans'' (Latin ''natans'' "swimming"), known solely from a single, near-complete fossil. ''Ambulocetus'' is among the best-studied of Eocene cetaceans, and serves as an instrumental find in the study of cetacean evolution and their transition from land to sea, as it was the first cetacean discovered to preserve a suite of adaptations consistent with an amphibious lifestyle. ''Ambulocetus'' is classified in the group Archaeoceti—the ancient forerunners of modern cetaceans whose members span the transition from land to sea—and in the family Ambulocetidae, which includes ''Himalayacetus'' and ''Gandakasia'' (also from the Eocene of the Indian subcontinent). ''Ambulocetus'' had a narrow, streamlined body, and a long, ...
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Pakicetid
Pakicetidae ("Pakistani whales") is an extinct family of Archaeoceti (early whales) that lived during the Early Eocene in Pakistan. Description described the first pakicetid, ''Ichthyolestes'', but at the time they did not recognize it as a cetacean, identifying it, instead, it as a fish-eating mesonychid. Robert West was the first to identify pakicetids as cetaceans in 1980 and, after discovering a braincase, Phillip Gingerich and Donald Russell described the genus ''Pakicetus'' in 1981. On October 26, 2016, a publication represented the idea that the emergence of cetacea in the Paleogene presents the best idea of microevolution that resulted in the phenotype of pakicetid. During the following two decades, more research resulted in additional pakicetid cranial material and by 2001 postcranial material for the family had been described. Though all parts of pakicetid postcrania are known, no complete skeleton from a single individual has been recovered. The pakicetid goldmine is t ...
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Protocetidae
Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America. Description There were many genera, and some of these are very well known (e.g., '' Rodhocetus''). Known protocetids had large fore- and hindlimbs that could support the body on land, and it is likely that they lived amphibiously: in the sea and on land. It is unclear at present whether protocetids had flukes (the horizontal tail fin of modern cetaceans). However, what is clear is that they are adapted even further to an aquatic life-style. In '' Rodhocetus'', for example, the sacrum – a bone that in land-mammals is a fusion of five vertebrae that connects the pelvis with the rest of the vertebral column – was divided into loose vertebrae. However, the pelvis retain a sacroiliac joint. Furthermore, the nasal openings are now halfway up the snout; a first step towards the telescoped condition in modern whales. ...
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Dorudon Atrox-lateral
''Dorudon'' ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside ''Basilosaurus'' 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with ''D. atrox'' measuring long and weighing . ''Dorudon'' lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand, and Western Sahara. Taxonomic history described ''Dorudon serratus'' based on a fragmentary maxilla and a few teeth found in South Carolina. He concluded that the teeth must have belonged to a mammal since they were two-rooted, that they must have been teeth from a juvenile since they were hollow, and also noted their similarity to the teeth then described for ''Zeuglodon'' (''Basilosaurus''). When exploring the type locality, Gibbes discovered a lower jaw and twelve caudal vertebrae, which he felt obliged to assign ...
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Remingtonocetidae
Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg. Description Remingtonocetids have long and narrow skulls with the external nare openings located on the front of the skull. Their frontal shields are narrow and their orbits small. Their mouth has a convex palate and an incompletely fused mandibular symphysis. The dental formula is . The anterior teeth are flattened mediolaterally, making them appear shark-like. In the postcranial skeleton, the cervical vertebrae are relatively long and the sacrum is composed of four vertebrae of which at least three are fused. The acetabular notch is narrow or closed and on the femoral head the fovea is absent. Cranial fossils are common but dental remains are rare. The postcrania morphology is based entirely on a single specimen of '' Kutchicetus'' which was small and had a long and muscular back and tail. Perhaps remintonocetids swam lik ...
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Rodhocetus
''Rodhocetus'' (from ''Rodho'', the geological anticline at the type locality, and ''cetus'', Latin for whale) is an extinct genus of protocetid early whale known from the Lutetian of Pakistan. The best-known protocetid, ''Rodhocetus'' is known from two partial skeletons that taken together give a complete image of an Eocene whale that had short limbs with long hands and feet that were probably webbed and a sacrum that was immobile with four partially fused sacral vertebrae. It is one of several extinct whale genera that possess land mammal characteristics, thus demonstrating the evolutionary transition from land to sea. Description left, Size of ''Rodhocetus'' relative to a human. ''Rodhocetus'' was a small whale measuring long. Throughout the 1990s, a close relationship between cetaceans and mesonychids, an extinct group of cursorial, wolf-like ungulates, was generally accepted based on morphological analyses. In the late 1990s, however, cladistic analyses based on molecul ...
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Evolution Of Cetaceans
The evolution of cetaceans is thought to have begun in the Indian subcontinent from even-toed ungulates 50 million years ago (mya) and to have proceeded over a period of at least 15 million years. Cetaceans are fully aquatic marine mammals belonging to the order Artiodactyla and branched off from other artiodactyls around 50  mya. Cetaceans are thought to have evolved during the Eocene (56-34 mya), the second epoch of the present-extending Cenozoic Era. Molecular and morphological analyses suggest Cetacea share a relatively recent closest common ancestor with hippopotami and that they are sister groups. Being mammals, they surface to breathe air; they have 5 finger bones (even-toed) in their fins; they nurse their young; and, despite their fully aquatic life style, they retain many skeletal features from their terrestrial ancestors.Thewissen, J. G. M., L. N. Cooper, J. C. George, and S. Bajpai. 2009. From land to water: the origin of whales, dolphins, and porpoises. Evolu ...
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Cetacea
Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver. While the majority of cetaceans live in marine environments, a small number exclusively reside in brackish water or fresh water. Having a cosmopolitan distribution, they can be found in some rivers and all of Earth's oceans, and many species inhabit vast ranges where they migrate with the changing of the seasons. Cetaceans are famous for their high intelligence and complex social behaviour as well as for the enormous size of some of the group's members, such as the blue whale which reaches a maximum confirmed length of 29.9 meters (98 feet) and a weight of 173 tonnes (190 short t ...
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Maiacetus
''Maiacetus'' ("mother whale") is a genus of early middle Eocene (c. 47.5 mya) cetacean from the Habib Rahi Formation of Pakistan. Paleobiology The genus contains a single species, ''Maiacetus inuus'', first described in 2009 on the basis of two specimens, including a specimen which has been interpreted as a pregnant female and its fetus. This represents the first description of a fetal skeleton of an archaeocete. The position of the fetus (head-first) suggests that the animal gave birth on land."Earliest whales gave birth on land"
'''', 3 February 2009
Modern whales generally give birth tail first, while land mammals give birth head first ...
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Himalayacetus
''Himalayacetus'' is an extinct genus of carnivorous aquatic mammal of the family Ambulocetidae. The holotype was found in Himachal Pradesh, India, (: paleocoordinates ) in what was the remnants of the ancient Tethys Ocean during the Early Eocene. This makes ''Himalayacetus'' the oldest archaeocete known, extending the fossil record of whales some 3.5 million years. ''Himalayacetus'' lived in the ancient coastline of the ancient Tethys Ocean before the Indian Plate had collided with the Cimmerian coast. Like ''Gandakasia'', ''Himalayacetus'' is only known from a single jaw fragment, making comparisons to other ambulocetids difficult. Description Upon its discovery, ''Himalayacetus'' was described as a pakicetid because the dentary has a small mandibular canal and a dentition similar to ''Pakicetus''. assigned ''Himalaycetus'' to the ambulocetids. Etymology ''Himalayacetus'' was named by . Its type is ''Himalayacetus subathuensis'' after the Himalayas, ''cetus'', "wh ...
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Cetacea
Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver. While the majority of cetaceans live in marine environments, a small number exclusively reside in brackish water or fresh water. Having a cosmopolitan distribution, they can be found in some rivers and all of Earth's oceans, and many species inhabit vast ranges where they migrate with the changing of the seasons. Cetaceans are famous for their high intelligence and complex social behaviour as well as for the enormous size of some of the group's members, such as the blue whale which reaches a maximum confirmed length of 29.9 meters (98 feet) and a weight of 173 tonnes (190 short t ...
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Pakicetus Fossil
''Pakicetus'' is an extinct genus of amphibious cetacean of the family Pakicetidae, which was endemic to Pakistan during the Eocene, about 50 million years ago. It was a wolf-like animal, about to long, and lived in and around water where it ate fish and small animals. The vast majority of paleontologists regard it as the most basal whale, representing a transitional stage between land mammals and whales. It belongs to the even-toed ungulates with the closest living non-cetacean relative being the hippopotamus. Description Based on the sizes of specimens, and to a lesser extent on composite skeletons, species of ''Pakicetus'' are thought to have been to in length. ''Pakicetus'' looked very different from modern cetaceans, and its body shape more resembled those of land-dwelling hoofed mammals. Unlike all later cetaceans, it had four fully functional long legs. ''Pakicetus'' had a long snout; a typical complement of teeth that included incisors, canines, premolars, and mol ...
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