Anomalocaridid
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Anomalocaridid
Radiodonta is an extinct order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa ''Anomalocaris canadensis'', ''Hurdia victoria'', ''Peytoia nathorsti'', '' Titanokorys gainessii, Cambroraster falcatus'' and '' Amplectobelua symbrachiata'', the Ordovician '' Aegirocassis benmoulai'' and the Devonian ''Schinderhannes bartelsi''. Etymology The name Radiodonta ( ...
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Anomalocarididae
Anomalocarididae (occasionally mis-spelt Anomalocaridae) is an extinct family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ... of Cambrian radiodonts, a group of stem-group arthropods. Around 1990s and early 2010s, Anomalocarididae included all radiodont species, hence the previous equivalent of the common name "anomalocaridid" to the whole Radiodonta. This is no longer the case after the revision done by Vinther et al. 2014, as Anomalocarididae restricted to only ''Anomalocaris'' and, if any, a few of closely-related genera since then. Wu et al. 2021 accepted only ''Anomalocaris'' (excluding ''"A." saron'', ''"A." kunmingensis'' and ''"A." briggsi'') and ''Lenisicaris'' as the member of Anomalocarididae, while ''Paranomalocaris'' is questionably included by some studies as well ...
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Dinocaridida
DinocarididaGreek, "Terrible crabs" – sometimes informally spelt Dinocarida, but the second 'id' is linguistically correct – see is a proposed fossil taxon of basal arthropods that flourished in the Cambrian period with occasional OrdovicianVan Roy, P.; Briggs, D. E. G. (2011). "A giant Ordovician anomalocaridid". Nature 473 (7348): 510–513. doi:10.1038/nature09920. edit and Devonian records. Characterized by a pair of frontal appendages and series of body flaps, the name of Dinocaridids comes from Greek, "deinos" and "caris" ("terrible crab"), referring to the suggested role of some of these members as the largest marine predators of their time. Dinocaridids are occasionally referred to as the 'AOPK group' by some literatures, as the group compose of Radiodonta (''Anomalocaris'' and relatives), Opabiniidae ('' Opabinia'' and relatives), ''Pambdelurion'' and ''Kerygmachela.'' It is most likely paraphyletic, with ''Kerygmachela'' and ''Pambdelurion'' more basal than the c ...
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Anomalocaris
''Anomalocaris'' ("unlike other shrimp", or "abnormal shrimp") is an extinct genus of radiodont, an order of early-diverging stem-group arthropods. The first fossils of ''Anomalocaris'' were discovered in the ''Ogygopsis'' Shale of the Stephen Formation in British Columbia, Canada by Joseph Frederick Whiteaves, with more examples found by Charles Doolittle Walcott in the Burgess Shale unit of the Stephen Formation. Other closely related fossils have been found in the older Emu Bay Shale of Australia, as well as possibly elsewhere. Originally several fossilized parts discovered separately (the mouth, frontal appendages and trunk) were thought to be three separate creatures, a misapprehension corrected by Harry B. Whittington and Derek Briggs in a 1985 journal article. With a body length close to 40 centimetres, ''A. canadensis'' is thought to be one of the earliest examples of an apex predator, though others have been found in older Cambrian lagerstätten deposits. Discovery ...
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Houcaris
''Houcaris'' is a genus of tamisiocarididid radiodonts known from Cambrian Series 2 of China and the United States. It contains two species, ''Houcaris saron'' and ''Houcaris magnabasis'', both of which were originally named as species of the related genus ''Anomalocaris''. The genus ''Houcaris'' was established for the two species in 2021 and honors Hou Xianguang, who had discovered and named the type species ''H. saron'' in 1995 along with his colleagues Jan Bergström and Per E. Ahlberg. Species ''Houcaris saron'' ''H. saron'', known from Maotianshan Shale in Yunnan, is first described in 1995 as ''Anomalocaris saron''. This species is only known from frontal appendages. There is a specimen (ELRC 20001) that is previously considered as whole body fossil of this species, but later study shows that this specimen is not belonging to this species, and later given own genus ''Innovatiocaris''. Length of frontal appendage is up to at least 12 cm. Sometimes considered to belo ...
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Cucumericrus
''Cucumericrus decoratus'' is a species of putative radiodont known from a few poorly preserved specimens. Only fragments of trunk cuticle and corresponded appendages had been revealed, while important radiodont features such as frontal appendages are unknown in this species. The trunk cuticle possess irregular wrinkles and may had been soft in life. Each of the trunk appendage compose of a dorsal flap-like element and a ventral stubby leg with unknown distal region, structurally comparable to the trunk appendages of gill lobopodians (dorsal flaps and ventral lobopods) and euarthropod biramous appendages (flap-like exopod and limb-like endopod). The legs have been interpreted as somewhere between annulated lobopod legs and segmented arthropod leg The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (mean ...
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Tamisiocarididae
Tamisiocarididae is a family of radiodonts, extinct marine animals related to arthropods, that bore finely-spined appendages that were presumably used in filter-feeding. When first discovered, the clade was named Cetiocaridae after a speculative evolution artwork, ''Bearded Ceticaris'' by John Meszaros, that depicted a hypothetical filter-feeding radiodont at a time before any were known to exist. However, the family name was not valid according to the International Code of Zoological Nomenclature, as no real genus named "Cetiocaris" exists, and in 2019 it was formally replaced by the name Tamisiocarididae, after the only valid genus of the clade at the time. The family is only known from Series 2 of the Cambrian, unlike other radiodont families, which persisted longer into the Cambrian. All known species would have lived in tropical or subtropical waters, suggesting a preference for warmer waters. Description Like most radiodonts, cetiocarids have spiny frontal appendages. Howe ...
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Laminacaris
''Laminacaris'' is a genus of extinct stem-group arthropods ( Radiodonta) that lived during the Cambrian period. It is monotypic with a single species ''Laminacaris chimera'', the fossil of which was described from the Chengjian biota of China in 2018. Around the same time, two specimens that were similar or of the same species were discovered at the Kinzers Formation in Pennsylvania, USA. The first specimens from China were three frontal appendages, without the other body parts. ''Laminacaris'' specimens were discovered from three sites in China: the mudstones of the Yu'anshan Member of the Chiungchussu Formation in eastern Yunnan Province, the Mafang section in the Haikou area at Kunming, and the Heimadi section, Chengjiang. The name is derived from Latin words, ''lamina'' meaning thin blade, and ''caris'' for crab; the species name refers to a Greek mythological creature, ''Chimera'', that has a body composed of parts of more than one animal. In 2022, it is treated that is po ...
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Innovatiocaris
''Innovatiocaris'' (meaning "innovation crab") is a genus of radiodont of uncertain family from the early Cambrian Chengjiang Lagerstätte of Yunnan Province, China. The genus contains two named species, ''I. maotianshanensis'', known from a nearly complete individual and isolated frontal appendages, and ''I.''? ''multispiniformis'', known from a complete frontal appendage. Discovery and naming The holotype specimen of ''Innovatiocaris maotianshanensis'', ELRC 20001 was long considered in scientific literature as "''Anomalocaris'' sp". or ''Anomalocaris saron'' (='' Houcaris saron''). However, in 2022, Zeng ''et al''. described it as a new genus and species of radiodont. The holotype consists of the nearly complete remains of a young individual, preserved on a part and counterpart. A brief description of the specimen was provided in Chen ''et al''. (1994), but, despite its popularity in popular science literature, it did not receiv a detailed description until 2022. ELRC 200 ...
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Hurdiidae
Hurdiidae is an extinct cosmopolitan family of radiodonts, a group of stem-group arthropods, which lived during the Paleozoic Era. It is the most long-lived radiodont clade, lasting from the Cambrian period to the Devonian period. Description Hurdiidae is characterized by frontal appendages with distal region composed of 5 subequal blade-like endites, alongside the enlarged head carapaces and tetraradial mouthpart ( oral cone). The frontal appendages of hurdiids have a distinctive morphology, with the appendage of most species bearing five equally-sized elongate blade-like ventral spines known as endites. Subsequent podomeres were reduced in size and with only small endites or none. Each podomere bore only a single endite, unlike other radiodonts, in which the endites were paired. In most species, the endites were curved medially, so that the appendages formed a basket-like structure. Some hurdiids had greater numbers of endites, with '' Cordaticaris'' bearing seven e ...
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Titanokorys Gainesii
''Titanokorys'' is a genus of extinct hurdiid radiodont (a grouping of primitive stem arthropods which lived during the early Paleozoic) that existed during the mid Cambrian. It is the largest member of its family from the Cambrian, with a body length of long, making it one of the largest animals of the time. It bears a resemblance to the related genus '' Cambroraster''. Fossils of ''T. gainesi'' were first found within Marble Canyon in 2018. The fossils were not named until 2021 because they were assumed to be giant specimens of ''Cambroraster''. The creature was one of several genera of radiodonts known from the Burgess Shale, with some of the others being ''Cambroraster'', ''Anomalocaris'', '' Peytoia'', and '' Hurdia''. ''Titanokorys'' is distinguished from other Burgess Shale radiodonts because of its large anterior sclerite (head covering carapace) and a pair of spines on the anteroventral sides. Based on the shape of its appendages, ''Titanokorys'' is speculated to ...
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Schinderhannes (genus)
''Schinderhannes bartelsi'' is a species of hurdiid radiodont (anomalocaridid) known from one specimen from the lower Devonian Hunsrück Slates. Its discovery was astonishing because previously, radiodonts were known only from exceptionally well-preserved fossil beds (Lagerstätten) from the Cambrian, 100 million years earlier. Discovery The single specimen was discovered in the Eschenbach-Bocksberg Quarry in Bundenbach, and is named after the outlaw Schinderhannes who frequented the area. Its specific epithet ''bartelsi'' honours Christoph Bartels, a Hunsrück Slate expert. The specimen is now housed in the Naturhistorisches Museum, Mainz. Morphology ''Schinderhannes'' is about long in full body length (6.8cm long excluding telson). Like other radiodonts, the head bears a pair of spiny frontal appendages, a radially-arranged ventral mouthpart ( oral cone), and a pair of large lateral compound eyes. Detailed morphology of the frontal appendages and oral cone are equivoca ...
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Hurdia
''Hurdia'' is an extinct genus of hurdiid radiodont that lived 505 million years ago during the Cambrian Period. As a radiodont like ''Peytoia'' and ''Anomalocaris'', it is part of the ancestral lineage that led to euarthropods. Description ''Hurdia'' was one of the largest organisms in the Cambrian oceans, ''H. victoria'' reached approximately in length, while ''H. triangulata'' reached up to just . Its head bore a pair of rake-like frontal appendages which shovelled food into its pineapple-ring-like mouth (oral cone). Like other hurdiids, ''Hurdia'' bore a large frontal carapace protruding from its head composed of three sclerites: a central component known as the H-element and two lateral components known as P-elements. The function of this organ remains mysterious; it cannot have been protective as there was no underlying soft tissue. Originally, it is estimated that body flaps ran along the sides of the organisms, from which large gills were suspended. However, anatomy o ...
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