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Ankylosaurus
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only ''A. magniventris''. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, ''Ankylosaurus'' is often considered the archetypal member of its group, despite having some unusual features. Possibly the largest-known ankylosaurid, ''Ankylosaurus'' is estimated to have been between long and to have weighed between . It was quadrupedal, with a broad, robust body. It had a wide, low skull, with two horns pointing backward from the ...
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Ankylosauridae
Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Paleogene extinction event. These animals were mainly herbivorous and were obligate quadrupeds, with leaf-shaped teeth and robust, scute-covered bodies. Ankylosaurids possess a distinctly domed and short snout, wedge-shaped osteoderms on their skull, scutes along their torso, and a tail club. Ankylosauridae is exclusively known from the northern hemisphere, with specimens found in western North America, Europe, and East Asia. The first discoveries within this family were of the genus ''Ankylosaurus'', by Peter Kaiser and Barnum Brown in Montana in 1906. Brown went on to name Ankylosauridae and the subfamily Ankylosaurinae in 1908. Anatomy Ankylosaurids are stout, solidly built, armoured dinosaurs. They possess accessory ossifications on ...
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Euoplocephalus
''Euoplocephalus'' ( ) is a genus of very large, herbivorous ankylosaurid dinosaurs, living during the Late Cretaceous of Canada. It has only one named species, ''Euoplocephalus tutus''. The first fossil of ''Euoplocephalus'' was found in 1897 in Alberta. In 1902, it was named ''Stereocephalus'', but that name had already been given to an insect, so it was changed in 1910. Later, many more ankylosaurid remains were found from the Campanian of North America and often made separate genera. In 1971, Walter Coombs concluded that they all belonged to ''Euoplocephalus'' which then would be one of the best-known dinosaurs. Recently however, experts have come to the opposite conclusion, limiting the authentic finds of ''Euoplocephalus'' to about a dozen specimens. These include a number of almost complete skeletons, so much is nevertheless known about the build of the animal. ''Euoplocephalus'' reached in length and in body mass. Its body was low-slung and very flat and wide, standin ...
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Ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous. Ankylosauria was first named by Henry Fairfield Osborn in 1923.Osborn, H. F. (1923). "Two Lower Cretaceous dinosaurs of Mongolia." ''American Museum Novitates'', 95: 1–1/ref> In the Linnaean classification system, the group is usually considered either a suborder or an infraorder. It is contained within the group Thyreophora, which also includes the stegosaurs, armored dinosaurs known for their combination of ...
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Thyreophora
Thyreophora ("shield bearers", often known simply as "armored dinosaurs") is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous. Thyreophorans are characterized by the presence of body armor lined up in longitudinal rows along the body. Primitive forms had simple, low, keeled scutes or osteoderms, whereas more derived forms developed more elaborate structures including spikes and plates. Most thyreophorans were herbivorous and had relatively small brains for their body size. Thyreophora includes various subgroups, including the suborders Ankylosauria and Stegosauria. In both the suborders, the forelimbs were much shorter than the hindlimbs, particularly in stegosaurs. The clade has been defined as the group consisting of all species more closely related to ''Ankylosaurus'' than to ''Triceratops''. Thyreophora is the sister group of Cerapoda within Genasauria. Groups of thyreophorans Basal thyreophorans Basal thyreo ...
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Club (anatomy)
In zoology, a club is a bony mass at the end of the tail of some dinosaurs and of some mammals, most notably the ankylosaurids and the glyptodonts, as well as meiolaniid turtles. It is thought that this was a form of defensive armour or weapon that was used to defend against predators, much in the same way as a thagomizer, possessed by stegosaurids, though at least in glyptodonts it is hypothesized it was used in fighting for mating rights. Among dinosaurs, the club was present mainly in ankylosaurids, although sauropods like ''Shunosaurus'' also possessed a tail club. The tail club is most often depicted on ''Ankylosaurus'', especially in encounters with larger predators such as ''Tyrannosaurus''. Victoria Arbour Victoria Megan Arbour is a Canadian evolutionary biologist and vertebrate palaeontologist at Royal BC Museum, where she is Curator of Palaeontology. An "expert on the armoured dinosaurs known as ankylosaurs", Arbour analyzes fossils and creates ... has established t ...
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Scollard Formation
The Scollard Formation is an Upper Cretaceous to lower Palaeocene stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. Its deposition spanned the time interval from latest Cretaceous to early Paleocene, and it includes sediments that were deposited before, during, and after the Cretaceous-Paleogene (K-Pg) extinction event. It is significant for its fossil record, and it includes the economically important coal deposits of the Ardley coal zone. Lithology The Scollard Formation consists primarily of sandstones and siltstones, interbedded with mudstones and, in the upper portion, coal seams, as well as minor amounts of bentonite. The sediments were eroded from the Canadian Cordillera, and were transported eastward by river systems and deposited in fluvial channel and floodplain environments. Stratigraphy The Scollard is usually classified as the uppermost formation of the Edmonton Group, although some early workers included it in the overlying ...
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Lance Formation
The Lance (Creek) Formation is a division of Late Cretaceous (dating to about 69 - 66 Ma) rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas. The Lance Formation is Late Maastrichtian in age (Lancian land mammal age), and shares much fauna with the Hell Creek Formation of Montana and North Dakota, the Frenchman Formation of southwest Saskatchewan, and the lower part of the Scollard Formation of Alberta. The Lance Formation occurs above the ''Baculites clinolobatus'' ammonite marine zone in Wyoming, the top of which has been dated to about 69 million years ago, and extends to the K-Pg boundary, 66 million years ago. However, the characteristic land vertebrate fauna of the Lancian age (which take its name from this formation) is only found in the upper strata of the Lance, roughly corresponding to the thinner equivalent formations such as the Hell C ...
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Nodosauridae
Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, were heavily armored dinosaurs adorned with rows of bony armor nodules and spines (osteoderms), which were covered in keratin sheaths. All nodosaurids, like other ankylosaurians, were medium-sized to large, heavily built, quadrupedal, herbivorous dinosaurs, possessing small, leaf-shaped teeth. Unlike ankylosaurids, nodosaurids lacked mace-like tail clubs, instead having flexible tail tips. Many nodosaurids had spikes projecting outward from their shoulders. One particularly well-preserved nodosaurid "mummy", known as the Suncor nodosaur (''Borealopelta markmitchelli''), preserved a nearly complete set of armor in life position, as well as the keratin covering and mineralized remains of the underlying skin, which indicate reddish dorsal pigme ...
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Ferris Formation
The Ferris Formation is a Late Cretaceous (~66 Ma) to Paleocene (66-63 Ma), fluvial-deltaic geological formation in southern Wyoming. It contains a variety of trace and body fossils, preserved in sandy fluvial channel deposits and overbank units. Dinosaur remains are fragmentary, but include ''Triceratops'', ''Tyrannosaurus'', dromaeosaurids, '' Paronychodon'', '' Ricardoestesia'', ''Edmontosaurus'', ''Edmontonia'', ''Ankylosaurus'', and ''Pachycephalosaurus''.Wroblewski, 1995 Some of the fluvial channels contain evidence of tidal influence and brackish water, in the form of tidal facies, shark teeth, and shrimp burrows. This demonstrates that the western shoreline of the Western Interior Sea was still within a few hundred kilometers even during the latest Cretaceous. The local K-T boundary is contained within a sandy channel deposit made up of stacked bars. Reworked Cretaceous fossils are preserved at the base of the channel complex, associated with mud rip-up clasts, and Paleo ...
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Tyrannosaurus
''Tyrannosaurus'' is a genus of large theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other tyrannosaurids. Fossils are found in a variety of rock formations dating to the Maastrichtian age of the Upper Cretaceous period, 68 to 66 million years ago. It was the last known member of the tyrannosaurids and among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the forelimbs of ''Tyrannosaurus'' were short but unusually powerful for their size, and they had two cla ...
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Anodontosaurus
''Anodontosaurus'' is an extinct genus of ankylosaurid dinosaurs within the subfamily Ankylosaurinae. It is known from the entire span of the Late Cretaceous Horseshoe Canyon Formation (mid Late Campanian to "middle" Maastrichtian stage, about 72.8-67 Ma ago) of southern Alberta, Canada. It contains two species, ''A. lambei'' and ''A. inceptus''. Discovery ''Anodontosaurus'' was named by Charles Mortram Sternberg in 1928, based on holotype CMN 8530, a partially preserved skeleton including the skull, half ring, armor and other postcranial remains.C. M. Sternberg (1929) "A toothless armoured dinosaur from the Upper Cretaceous of Alberta." ''Canada Department of Mines Geological Survey Bulletin (Geological Series)'' 54(49):28-33 The badly crushed skeleton was collected by Sternberg in 1916 from a Canadian Museum of Nature quarry, 8 miles southwest of Morrin. It was collected from the upper part of the Lower Horseshoe Canyon Formation (unit 2), dating to the latest Campan ...
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Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of Montana, North Dakota, South Dakota, and Wyoming. In Montana, the Hell Creek Formation overlies the Fox Hills Formation. The site of Pompeys Pillar National Monument is a small isolated section of the Hell Creek Formation. In 1966, the Hell Creek Fossil Area was designated as a National Natural Landmark by the National Park Service. It is a series of fresh and brackish-water clays, mudstones, and sandstones deposited during the Maastrichtian and Danian (respectively, the end of the Cretaceous period and the beginning of the Paleogene) by fluvial activity in fluctuating river channels and deltas and very occasional peaty swamp deposits along the low-lying eastern continental margin fronting the late Cretaceous Western Interior Seaway. Th ...
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