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The wildcat is a species complex comprising two small wild cat
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
: the
European wildcat The European wildcat (''Felis silvestris'') is a small wildcat species native to continental Europe, Scotland, Turkey and the Caucasus. It inhabits forests from the Iberian Peninsula, Italy, Central and Eastern Europe to the Caucasus. Its ...
(''Felis silvestris'') and the African wildcat (''F. lybica''). The European wildcat inhabits
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
and the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
, while the African wildcat inhabits semi-
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ...
landscapes and
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate gras ...
s in
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...
,
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the fo ...
, into western
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
and western
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
. The wildcat species differ in fur pattern, tail, and size: the European wildcat has long fur and a bushy tail with a rounded tip; the smaller African wildcat is more faintly striped, has short sandy-gray fur and a tapering tail; the Asiatic wildcat (''F. lybica ornata'') is spotted. The wildcat and the other members of the cat family had a common ancestor about 10–15 million years ago. The European wildcat
evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variati ...
during the Cromerian Stage about 866,000 to 478,000 years ago; its direct ancestor was '' Felis lunensis''. The ''silvestris'' and ''lybica'' lineages probably diverged about 173,000 years ago. The wildcat has been categorized as
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
since 2002, since it is widely distributed, and the global population is considered stable and exceeding 20,000 mature individuals. However, in some range countries both wildcat species are considered threatened by introgressive hybridisation with the domestic cat (''F. catus'') and transmission of diseases. Localized threats include being hit by vehicles, and persecution. The association of African wildcats and humans appears to have developed along with the establishment of settlements during the Neolithic Revolution, when
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s in grain stores of early
farmer A farmer is a person engaged in agriculture, raising living organisms for food or raw materials. The term usually applies to people who do some combination of raising field crops, orchards, vineyards, poultry, or other livestock. A farmer m ...
s attracted wildcats. This association ultimately led to it being
tamed A tame animal is an animal that is relatively tolerant of human presence. Tameness may arise naturally (as in the case, for example, of island tameness) or due to the deliberate, human-directed process of training an animal against its initially ...
and domesticated: the domestic cat is the direct descendant of the African wildcat. It was one of the revered cats in ancient Egypt. The European wildcat has been the subject of
mythology Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narra ...
and
literature Literature is any collection of Writing, written work, but it is also used more narrowly for writings specifically considered to be an art form, especially prose fiction, drama, and poetry. In recent centuries, the definition has expanded to ...
.


Taxonomy

''Felis (catus) silvestris'' was the scientific name used in 1777 by Johann von Schreber when he described the European wildcat based on descriptions and names proposed by earlier naturalists such as
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...
, Ulisse Aldrovandi and Conrad Gessner. ''Felis lybica'' was the name proposed in 1780 by Georg Forster, who described an African wildcat from Gafsa on the Barbary Coast. In subsequent decades, several naturalists and explorers described 40 wildcat
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...
s collected in European, African and Asian range countries. In the 1940s, the taxonomist Reginald Innes Pocock reviewed the collection of wildcat skins and skulls in the
Natural History Museum, London The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum an ...
, and designated seven ''F. silvestris'' subspecies from Europe to
Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
, and 25 ''F. lybica'' subspecies from
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and West to
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the fo ...
. Pocock differentiated the: * Forest wildcat subspecies (''silvestris'' group) * Steppe wildcat subspecies (''ornata''-''caudata'' group): is distinguished from the forest wildcat by being smaller, with comparatively lighter fur colour, and longer and more sharply-pointed tails. The domestic cat is thought to have derived from this group. * Bush wildcat subspecies (''ornata''-''lybica'' group): is distinguished from the steppe wildcat by paler fur, well-developed spot patterns and bands. In 2005, 22 subspecies were recognized by the authors of '' Mammal Species of the World'', who allocated subspecies largely in line with Pocock's assessment. In 2017, the Cat Classification Task Force revised the taxonomy of the Felidae, and recognized the following as valid taxa:


Evolution

The wildcat is a member of the Felidae, a family that had a common ancestor about 10–15 million years ago. ''Felis'' species diverged from the Felidae around 6–7 million years ago. The European wildcat diverged from ''Felis'' about 1.09 to 1.4 million years ago. The European wildcat's direct ancestor was '' Felis lunensis'', which lived in Europe in the late
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Villafranchian Villafranchian age ( ) is a period of geologic time (3.5–1.0 Ma) spanning the Late Pliocene and Early Pleistocene used more specifically with European Land Mammal Ages. Named by Italian geologist Lorenzo Pareto for a sequence of terrestrial se ...
periods.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
remains indicate that the transition from ''lunensis'' to ''silvestris'' was completed by the Holstein interglacial about 340,000 to 325,000 years ago. Craniological differences between the European and African wildcats indicate that the wildcat probably migrated during the Late Pleistocene from Europe into the Middle East, giving rise to the steppe wildcat
phenotype In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology (biology), morphology or physical form and structure, its Developmental biology, developmental proc ...
.
Phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...
research revealed that the ''lybica'' lineage probably diverged from the ''silvestris'' lineage about 173,000 years ago.


Characteristics

The wildcat has pointed ears, which are moderate in length and broad at the base. Its whiskers are white, number 7 to 16 on each side and reach in length on the muzzle. Whiskers are also present on the inner surface of the paw and measure . Its eyes are large, with vertical pupils and yellowish-green irises. The eyelashes range from in length, and can number six to eight per side. The European wildcat has a greater
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
volume than the domestic cat, a ratio known as
Schauenberg's index Schauenberg's index is the ratio of skull length to cranial capacity. This index was introduced by Paul Schauenberg in 1969 as a method to identify European wildcat (''Felis silvestris'') skulls and distinguish them from domestic cat The c ...
. Further, its skull is more spherical in shape than that of the jungle cat (''F. chaus'') and
leopard cat The leopard cat (''Prionailurus bengalensis'') is a small wild cat native to continental South, Southeast, and East Asia. Since 2002 it has been listed as Least Concern on the IUCN Red List as it is widely distributed although threatened by hab ...
(''Prionailurus bengalensis''). Its
dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
is relatively smaller and weaker than the jungle cat's. Both wildcat species are larger than the domestic cat. The European wildcat has relatively longer legs and a more robust build compared to the domestic cat. The tail is long, and usually slightly exceeds one-half of the animal's body length. The species size varies according to Bergmann's rule, with the largest specimens occurring in cool, northern areas of Europe and Asia such as
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
,
Manchuria Manchuria is an exonym (derived from the endo demonym " Manchu") for a historical and geographic region in Northeast Asia encompassing the entirety of present-day Northeast China (Inner Manchuria) and parts of the Russian Far East (Outer M ...
and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
. Males measure in head to body length, in tail length, and normally weigh . Females are slightly smaller, measuring in body length and in tail length, and weighing . Both sexes have two thoracic and two
abdominal The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the to ...
teats. Both sexes have pre-
anal gland Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving ...
s, consisting of moderately sized sweat and
sebaceous gland A sebaceous gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in the greatest n ...
s around the anal opening. Large-sized sebaceous and scent glands extend along the full length of the tail on the dorsal side. Male wildcats have pre-anal pockets on the tail, activated upon reaching sexual maturity, play a significant role in reproduction and territorial marking.


Distribution and habitat

The European wildcat inhabits
temperate broadleaf and mixed forest Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. These ...
s in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
and the Caucasus. In the
Iberian peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
, it occurs from sea level to in the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
. Between the late 17th and mid 20th centuries, its European range became fragmented due to large-scale hunting and regional extirpation. It is possibly extinct in the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
, and considered regionally extinct in
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
, though vagrants from
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
are spreading into Austria. It has never inhabited Fennoscandia or
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, an ...
.
Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...
is the only island in the
Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
with a native wildcat population. The African wildcat lives in a wide range of habitats except rainforest, but throughout the savannahs of Africa from Mauritania on the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
coast eastwards to the Horn of Africa up to altitudes of . Small populations live in the Sahara and
Nubian Desert The Nubian Desert ( ar, صحراء النوبة, ''Şaḩrā’ an Nūbyah'') is in the eastern region of the Sahara Desert, spanning approximately 400,000 km2 of northeastern Sudan and northern Eritrea, between the Nile and the Red Sea ...
s, Karoo region, Kalahari and Namib Deserts. It occurs around the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...
's periphery to the Caspian Sea, encompassing
Mesopotamia Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
and Palestine region. In Central Asia, it ranges into
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwes ...
and southern
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
, and in
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;; ...
into the
Thar Desert The Thar Desert, also known as the Great Indian Desert, is an arid region in the north-western part of the Subcontinent that covers an area of and forms a natural boundary between India and Pakistan. It is the world's 20th-largest desert, ...
and arid regions in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
.


Behaviour and ecology

Both wildcat species are largely nocturnal and
solitary Solitary is the state of being alone or in solitude. The term may refer to: * shortened form of solitary confinement * Solitary animal, an animal that does not live with others in its species * Solitary but social, a type of social organization ...
, except during the breeding period and when females have young. The size of home ranges of females and males varies according to terrain, the availability of food, habitat quality and the age structure of the population. Male and female home ranges overlap, though core areas within
territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
are avoided by other cats. Females tend to be more sedentary than males, as they require an exclusive hunting area when raising kittens. Wildcats usually spend the day in a hollow tree, a rock crevice or in dense thickets. It is also reported to shelter in abandoned burrows of other species such as of red fox (''Vulpes vulpes'') and in European badger (''Meles meles'') setts in Europe, and of fennec (''Vulpes zerda'') in Africa. When threatened, it retreats into a burrow, rather than climb trees. When taking residence in a tree hollow, it selects one low to the ground. Dens in rocks or burrows are lined with dry grasses and bird feathers. Dens in tree hollows usually contain enough sawdust to make lining unnecessary. If the den becomes infested with
flea Flea, the common name for the order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by ingesting the blood of their hosts. Adult fleas grow to about long, ...
s, the wildcat shifts to another den. During winter, when snowfall prevents the European wildcat from travelling long distances, it remains within its den until travel conditions improve. Territorial marking consists of spraying urine on trees, vegetation and rocks, depositing faeces in conspicuous places, and leaving scent marks through glands in its paws. It also leaves visual marks by scratching trees.


Hunting and prey

Sight and hearing are the wildcat's primary senses when hunting. It lies in wait for prey, then catches it by executing a few leaps, which can span three metres. When hunting near water courses, it waits on trees overhanging the water. It kills small prey by grabbing it in its claws, and piercing the neck or occiput with its fangs. When attacking large prey, it leaps upon the animal's back, and attempts to bite the neck or
carotid In anatomy, the left and right common carotid arteries (carotids) (Entry "carotid"
in
The European wildcat primarily preys on small mammals such as European rabbit (''Oryctolagus cuniculus'') and
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s. It also preys on dormice, hares, nutria (''Myocastor coypus'') and
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, especially
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form ...
s and other
waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which ...
,
galliformes Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
, pigeons and passerines. It can consume large
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
fragments. Although it kills insectivores such as
moles Moles can refer to: * Moles de Xert, a mountain range in the Baix Maestrat comarca, Valencian Community, Spain *The Moles (Australian band) *The Moles, alter ego of Scottish band Simon Dupree and the Big Sound People * Abraham Moles, French engin ...
and
shrew Shrews (family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to differ ...
s, it rarely eats them. When living close to human settlements, it preys on
poultry Poultry () are domesticated birds kept by humans for their eggs, their meat or their feathers. These birds are most typically members of the superorder Galloanserae (fowl), especially the order Galliformes (which includes chickens, qu ...
. In the wild, it consumes up to of food daily. The African wildcat preys foremost on murids, to a lesser extent also on birds, small reptiles and
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
s.


Reproduction and development

The wildcat has two estrus periods, one in December–February and another in May–July. Estrus lasts 5–9 days, with a
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once ...
lasting 60–68 days. Ovulation is induced through copulation. Spermatogenesis occurs throughout the year. During the mating season, males fight viciously, and may congregate around a single female. There are records of male and female wildcats becoming temporarily monogamous. Kittens are usually born between April and May, and up to August. Litter size ranges from 1–7 kittens. Kittens are born with closed eyes and are covered in a fuzzy coat. They weigh at birth, and kittens under usually do not survive. They are born with pink paw pads, which blacken at the age of three months, and blue eyes, which turn amber after five months. Their eyes open after 9–12 days, and their incisors erupt after 14–30 days. The kittens' milk teeth are replaced by their permanent dentition at the age of 160–240 days. The kittens start hunting with their mother at the age of 60 days, and start moving independently after 140–150 days.
Lactation Lactation describes the secretion of milk from the mammary glands and the period of time that a mother lactates to feed her young. The process naturally occurs with all sexually mature female mammals, although it may predate mammals. The proces ...
lasts 3–4 months, though the kittens eat meat as early as 1.5 months of age. Sexual maturity is attained at the age of 300 days. Similarly to the domestic cat, the physical development of African wildcat kittens over the first two weeks of their lives is much faster than that of European wildcats. The kittens are largely fully grown by 10 months, though skeletal growth continues for over 18–19 months. The family dissolves after roughly five months, and the kittens disperse to establish their own territories. Their maximum life span is 21 years, though they usually live up to 13–14 years. Generation length of the wildcat is about eight years.


Predators and competitors

Because of its habit of living in areas with rocks and tall trees for refuge, dense thickets and abandoned burrows, wildcats have few natural predators. In Central Europe, many kittens are killed by European pine marten (''Martes martes''), and there is at least one account of an adult wildcat being killed and eaten. Competitors include the golden jackal (''Canis aureus''), red fox, marten, and other predators. In the steppe regions of Europe and Asia, village dogs constitute serious enemies of wildcats, along with the much larger Eurasian lynx, one of the rare habitual predators of healthy adult wildcats. In Tajikistan, the
grey wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly u ...
(''Canis lupus'') is the most serious competitor, having been observed to destroy cat burrows. Birds of prey, including Eurasian eagle-owl (''Bubo bubo'') and saker falcon (''Falco cherrug''), have been recorded to kill wildcat kittens.
Golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird ...
(''Aquila chrysaetos'') are known to hunt both adults and kittens.
Seton Gordon Seton Gordon CBE (1886–1977) was a Scottish naturalist, photographer and folklorist. Gordon began exploring the Highlands of Scotland as a boy, particularly the Cairngorms. He later became a world-famous naturalist, photographer and folklori ...
recorded an instance where a wildcat fought a golden eagle, resulting in the deaths of both combatants. In Africa, wildcats are occasionally killed and eaten by African rock python (''Python sebae'') and martial eagle (''Polemaetus bellicosus'').


Threats

Wildcat populations are foremost threatened by hybridization with the domestic cat. Mortality due to traffic accidents is a threat especially in Europe. The wildcat population in Scotland has declined since the turn of the 20th century due to
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
and persecution by landowners. In the former Soviet Union, wildcats were caught accidentally in traps set for European pine marten. In modern times, they are caught in unbaited traps on pathways or at abandoned trails of red fox, European badger, European hare or pheasant. One method of catching wildcats consists of using a modified muskrat trap with a spring placed in a concealed pit. A scent trail of pheasant viscera leads the cat to the pit. Wildcat skins were of little commercial value and sometimes converted into imitation sealskin; the fur usually fetched between 50 and 60 kopecks. Wildcat skins were almost solely used for making cheap
scarf A scarf, plural ''scarves'', is a piece of fabric worn around the neck or head for warmth, sun protection, cleanliness, fashion, religious reasons, or used to show the support for a sports club or team. They can be made in a variety of diff ...
s, muffs and coats for ladies.


Conservation

Wildcat species are protected in most range countries and listed in CITES Appendix II. The European wildcat is also listed in Appendix II of the Berne Convention on the Conservation of European Wildlife and Natural Habitats and in the
European Union The European Union (EU) is a supranational union, supranational political union, political and economic union of Member state of the European Union, member states that are located primarily in Europe, Europe. The union has a total area of ...
's Habitats and Species Directive. Conservation Action Plans have been developed in Germany and Scotland.


In culture


Domestication

An African wildcat skeleton excavated in a 9,500-year-old Neolithic grave in Cyprus is the earliest known indication for a close relationship between a human and a possibly tamed cat. As no cat species is native to Cyprus, this discovery indicates that Neolithic farmers may have brought cats to Cyprus from the Near East. Results of
genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar work ...
and morphological research corroborated that the African wildcat is the ancestor of the domestic cat. The first individuals were probably domesticated in the
Fertile Crescent The Fertile Crescent ( ar, الهلال الخصيب) is a crescent-shaped region in the Middle East, spanning modern-day Iraq, Syria, Lebanon, Israel, Palestine and Jordan, together with the northern region of Kuwait, southeastern region of ...
around the time of the introduction of agriculture.
Mural A mural is any piece of graphic artwork that is painted or applied directly to a wall, ceiling or other permanent substrate. Mural techniques include fresco, mosaic, graffiti and marouflage. Word mural in art The word ''mural'' is a Spanis ...
s and statuettes depicting cats as well mummified cats indicate that it was commonly kept by ancient Egyptians since at least the Twelfth Dynasty of Egypt.


In mythology

Fables of the Cat Sìth, a fairy creature described as resembling a large white-chested black cat, are thought to have been inspired by the Kellas cat, itself thought to be a free-ranging crossbreed between a European wildcat and a domestic cat. In 1693,
William Salmon William Salmon (1644–1713) was an English empiric doctor and a writer of medical texts. He advertised himself as a "Professor of Physick". Salmon held an equivocal place in the medical community. He led apothecaries in opposing attempts by ...
mentioned how body parts of the wildcat were used for medicinal purposes; its flesh for treating
gout Gout ( ) is a form of inflammatory arthritis characterized by recurrent attacks of a red, tender, hot and swollen joint, caused by deposition of monosodium urate monohydrate crystals. Pain typically comes on rapidly, reaching maximal intens ...
, its fat for dissolving
tumour A neoplasm () is a type of abnormal and excessive growth of tissue. The process that occurs to form or produce a neoplasm is called neoplasia. The growth of a neoplasm is uncoordinated with that of the normal surrounding tissue, and persists ...
s and easing pain, its blood for curing " falling sickness", and its excrement for treating baldness.


In heraldry

The
Picts The Picts were a group of peoples who lived in what is now northern and eastern Scotland (north of the Firth of Forth) during Late Antiquity and the Early Middle Ages. Where they lived and what their culture was like can be inferred from ea ...
venerated wildcats, having probably named Caithness (Land of the Cats) after them. According to the foundation myth of the Catti tribe, their ancestors were attacked by wildcats upon landing in Scotland. Their ferocity impressed the Catti so much, that the wildcat became their symbol. The progenitors of Clan Sutherland use the wildcat as symbol on their family crest. The clan's chief bears the title ''Morair Chat'' (Great Man of the Cats). The wildcat is considered an
icon An icon () is a religious work of art, most commonly a painting, in the cultures of the Eastern Orthodox, Oriental Orthodox, and Catholic churches. They are not simply artworks; "an icon is a sacred image used in religious devotion". The mos ...
of Scottish wilderness, and has been used in clan heraldry since the 13th century. The Clan Chattan Association (also known as the Clan of Cats) comprises 12 clans, the majority of which display the wildcat on their badges.


In literature

Shakespeare William Shakespeare ( 26 April 1564 – 23 April 1616) was an English playwright, poet and actor. He is widely regarded as the greatest writer in the English language and the world's pre-eminent dramatist. He is often called England's nation ...
referenced the wildcat three times:


References


Further reading

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External links

* * * * * * * * Scottish wildcat ** ** {{Authority control Felis Mammals described in 1777 Mammals of Africa Mammals of Asia Mammals of Europe Mammals of Azerbaijan Mammals of Pakistan Mammals of the Middle East Mammals of Uganda