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The weka, also known as the Māori hen or woodhen (''Gallirallus australis'') is a
flightless bird Flightless birds are birds that through evolution lost the ability to fly. There are over 60 extant species, including the well known ratites (ostriches, emu, cassowaries, rheas, and kiwi) and penguins. The smallest flightless bird is the I ...
species of the
rail Rail or rails may refer to: Rail transport *Rail transport and related matters * Rail (rail transport) or railway lines, the running surface of a railway Arts and media Film * ''Rails'' (film), a 1929 Italian film by Mario Camerini * ''Rail'' ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island country ...
. It is the only extant member of the genus '' Gallirallus''. Four
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized but only two (northern/southern) are supported by genetic evidence. The weka are sturdy brown birds, about the size of a chicken. As
omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
s, they feed mainly on
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s and fruit. Weka usually lay
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
between August and January; both sexes help to incubate.


Description

Weka are large rails. They are predominantly rich brown mottled with black and grey; the brown shade varies from pale to dark depending on subspecies. The male is the larger sex at in length and in weight. Females measure in length and weigh . The reduced wingspan ranges from . The relatively large, reddish-brown beak is about long, stout and tapered, and used as a weapon. The pointed tail is near-constantly being flicked, a sign of unease characteristic of the rail family. Weka have sturdy legs and reduced wings.


Taxonomy and distribution

The common name, "weka", is a Māori word. The species was named ''Rallus australis'' by
Anders Erikson Sparrman Anders Sparrman (27 February 1748, Tensta, Uppland – 9 August 1820) was a Swedish naturalist, abolitionist and an apostle of Carl Linnaeus. Biography left, Miniature of Sparrman at the time of his travels with James Cook. By unknown arti ...
in 1789. Sparrman published the information in ''Museum Carlsonianum'', four fascicules based on specimens collected while voyaging with
Captain James Cook James Cook (7 November 1728 Old Style date: 27 October – 14 February 1779) was a British explorer, navigator, cartographer, and captain in the British Royal Navy, famous for his three voyages between 1768 and 1779 in the Pacific Ocean and ...
between 1772 and 1775. ''Australis'' is Latin for "southern".
Johann Georg Wagler Johann Georg Wagler (28 March 1800 – 23 August 1832) was a German herpetologist and ornithologist. Wagler was assistant to Johann Baptist von Spix, and gave lectures in zoology at the Ludwig Maximilian University of Munich after it was moved ...
's suggestion of the genus '' Ocydromus'' in 1830 to describe each weka as a species was generally adopted. However, weka were later classified as a single species in the genus '' Gallirallus'' with four subspecies. The buff weka (''Gallirallus australis hectori'') formerly inhabited the eastern districts of the
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman S ...
but is now confined to
Chatham Island Chatham Island ( ) (Moriori: ''Rēkohu'', 'Misty Sun'; mi, Wharekauri) is by far the largest island of the Chatham Islands group, in the south Pacific Ocean off the eastern coast of New Zealand's South Island. It is said to be "halfway bet ...
and
Pitt Island Pitt Island is the second largest island in the Chatham Archipelago, New Zealand. It is called ''Rangiauria'' in Māori and ''Rangiaotea'' in ''Moriori.Government of New Zealand, Dept. of Conservation (1999) Chatham IslandsConservation Manage ...
to which it was introduced in the early 1900s, and where they are widely hunted and eaten in the autumn (accepted because they are introduced). Reintroduction into
Canterbury Canterbury (, ) is a cathedral city and UNESCO World Heritage Site, situated in the heart of the City of Canterbury local government district of Kent, England. It lies on the River Stour. The Archbishop of Canterbury is the primate of the ...
has been unsuccessful so far but introductions to Mao Waho Island (in
Lake Wanaka A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much larger ...
in 2004) and from there to
Pigeon Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
and Pig Island (in
Lake Wakatipu Lake Wakatipu ( mi, Whakatipu Waimāori) is an inland lake (finger lake) in the South Island of New Zealand. It is in the southwest corner of the Otago region, near its boundary with Southland. ''Lake Wakatipu'' comes from the original Māori n ...
in 2005/2006) have been much more successful. It has a lighter overall colouring than the other subspecies. The North Island weka (''Gallirallus australis greyi'') is represented by original populations in Northland and
Poverty Bay Poverty Bay (Māori: ''Tūranganui-a-Kiwa'') is the largest of several small bays on the east coast of New Zealand's North Island to the north of Hawke Bay. It stretches for from Young Nick's Head in the southwest to Tuaheni Point in the nor ...
, and by liberations elsewhere from that stock. This subspecies differs in its greyer underparts, and brown rather than reddish coloured legs.Penguin Pocket Guides: New Zealand's Native Birds of Bush and Countryside. Pages 76-77. 1996. Published by Penguin Books. The western weka (''Gallirallus australis australis'') is found mainly in the northern and western regions of the South Island from
Nelson Nelson may refer to: Arts and entertainment * ''Nelson'' (1918 film), a historical film directed by Maurice Elvey * ''Nelson'' (1926 film), a historical film directed by Walter Summers * ''Nelson'' (opera), an opera by Lennox Berkeley to a lib ...
to
Fiordland Fiordland is a geographical region of New Zealand in the south-western corner of the South Island, comprising the westernmost third of Southland. Most of Fiordland is dominated by the steep sides of the snow-capped Southern Alps, deep lake ...
. Distinguished by dark red-brown and black streaking on the breast, the western weka has two distinct colour phases, that of the southernmost range showing a greater degree of black. The Stewart Island weka (''Gallirallus australis scotti'') is smaller than the other subspecies and, like the western weka, has two colour phases; a chestnut form – similar to the chestnut-phase western weka – and a black phase which is not as dark as the black western weka. Reintroduced populations are confined to
Stewart Island/Rakiura Stewart Island ( mi, Rakiura, ' glowing skies', officially Stewart Island / Rakiura) is New Zealand's third-largest island, located south of the South Island, across the Foveaux Strait. It is a roughly triangular island with a total land ar ...
and outliers, and to
Kapiti Island Kapiti Island () is an island about off the west coast of the lower North Island of New Zealand. It is long, running southwest/northeast, and roughly wide, being more or less rectangular in shape, and has an area of . Its name has been used s ...
to which it was introduced. Weka are host to two species of feather lice (''Rallicola harrisoni'' and ''Pseudomenopon pilgrimi'') which show the same north–south population structure seen in their hosts.


Behaviour


Habitat and diet

Weka occupy areas such as forests,
sub-alpine Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
grassland,
sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...
s,
rocky shore A rocky shore is an intertidal area of seacoasts where solid rock predominates. Rocky shores are biologically rich environments, and are a useful "natural laboratory" for studying intertidal ecology and other biological processes. Due to their ...
s and modified semi-urban environments. They are omnivorous, with a diet comprising 30% animal foods and 70% plant foods. Animal foods include
earthworm An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. Th ...
s,
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
e, beetles,
wētā Wētā (also spelt weta) is the common name for a group of about 100 insect species in the families Anostostomatidae and Rhaphidophoridae endemic to New Zealand. They are giant flightless crickets, and some are among the heaviest insects in ...
, ants, grass grubs, slugs, snails, insect eggs, slaters, frogs, spiders, rats, mice, and small birds. Stewart Island weka (''G. a. scotti'') have been observed preying on sooty shearwater (''Puffinus griseus'') eggs and chicks. Plant foods include leaves, grass, berries and seeds. Weka are important in the bush as seed dispersers, distributing seeds too large for smaller berry-eating birds. Where the weka is relatively common, their furtive curiosity leads them to search around houses and camps for food scraps, or anything unfamiliar and transportable. They have been known to take shiny objects in particular.


Breeding and nesting

The breeding season varies, but when food is plentiful, weka can raise up to four broods throughout the whole year.
Nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
s are made on the ground under the cover of thick vegetation, and built by making grass (or similar material) into a bowl to hold about four eggs. On average, female weka lay three creamy or pinkish eggs blotched with brown and mauve. Both sexes incubate. The chicks hatch after a month, and are fed by both parents until fully grown between six and ten weeks.


Conservation status

Weka are classed as a
vulnerable species A vulnerable species is a species which has been categorized by the International Union for Conservation of Nature as being threatened with extinction unless the circumstances that are threatening its survival and reproduction improve. Vulnerab ...
. The Department of Conservation's weka recovery plan, approved in 1999, aims to improve the conservation status of threatened weka, clarify the status of data deficient weka, maintain the non-threatened status of other weka, and eventually restore all weka to their traditional ranges as a significant component of the ecosystems. Weka are problematic in conservation; some subspecies are threatened, but have been a problem to other threatened wildlife on offshore islands, especially when introduced to an island that they would not naturally inhabit. Weka are unable to withstand the current pressures faced in both the
North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-largest ...
and
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman S ...
. However, they can be very productive in good conditions and high food availability. Year-round breeding has been recorded at several sites with up to 14 young produced in a year. Weka populations can persist in highly modified habitats, but they have disappeared from huge areas of their former range, suggesting that they can adapt to a wide range of conditions but are particularly vulnerable to threats.


Threats

The Department of Conservation identifies eight main threats to weka.
Predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
by
ferret The ferret (''Mustela furo'') is a small, domesticated species belonging to the family Mustelidae. The ferret is most likely a domesticated form of the wild European polecat (''Mustela putorius''), evidenced by their interfertility. Other must ...
s, cats, and dogs are a threat to adult weka;
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
s and ferrets are a threat to chicks; stoats and rats are a threat to eggs. It faces competition with introduced species for fruits and invertebrates, and suffers from the impact browsers have on forest composition and regeneration. Habitat depletion is caused by the modification and degradation of forests and wetlands. Diseases and parasites have been associated with population declines, although little is known. Drought has been implicated in the disappearance of weka from some areas. In some regions, motor vehicles cause a significant amount of
roadkill Roadkill is an animal or animals that have been struck and killed by drivers of motor vehicles on highways. Wildlife-vehicle collisions (WVC) have increasingly been the topic of academic research to understand the causes, and how it can be mi ...
death. Pest control operations sometimes kill weka, as they have ground foraging habits vulnerable to poison baits, and traps are laid in a way that weka can reach.
Genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species, it ranges widely from the number of species to differences within species and can be attributed to the span of survival for a species. It is dis ...
can be lost during the transmission of genes through generations, affecting isolated populations.


Human interaction

Weka are significant to some Māori
iwi Iwi () are the largest social units in New Zealand Māori society. In Māori roughly means "people" or "nation", and is often translated as "tribe", or "a confederation of tribes". The word is both singular and plural in the Māori language, an ...
(tribes) who admire their curiosity and feisty, bold personality traits which have led to them being relatively easy to catch. Weka were used by the Māori as a source of food, perfume, oil to treat
inflammation Inflammation (from la, inflammatio) is part of the complex biological response of body tissues to harmful stimuli, such as pathogens, damaged cells, or irritants, and is a protective response involving immune cells, blood vessels, and molec ...
s, feathers in clothing and lures to catch dogs. Early European explorers and settlers frequently encountered and utilised weka; they were called "bush hens".


References


External links


BirdLife Species Factsheet.
* {{Taxonbar, from=Q756714 Endemic birds of New Zealand Flightless birds Gallirallus Birds described in 1786 Taxa named by Anders Sparrman