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The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', its former genus name) is one of the two wild
South American South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
camelid Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
s, which live in the high
alpine Alpine may refer to any mountainous region. It may also refer to: Places Europe * Alps, a European mountain range ** Alpine states, which overlap with the European range Australia * Alpine, New South Wales, a Northern Village * Alpine National P ...
areas of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
, the other being the
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco ...
, which lives at lower elevations. Vicuñas are relatives of the
llama The llama (; ) (''Lama glama'') is a domesticated South American camelid, widely used as a meat and pack animal by Andean cultures since the Pre-Columbian era. Llamas are social animals and live with others as a herd. Their wool is soft ...
, and are now believed to be the wild ancestor of domesticated
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. It is similar to, and often confused with, the llama. However, alpacas are often noticeably smaller than llamas. The two animals are closely related and can success ...
s, which are raised for their coats. Vicuñas produce small amounts of extremely fine
wool Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool. ...
, which is very expensive because the animal can only be
shorn Sheep shearing is the process by which the woollen fleece of a sheep is cut off. The person who removes the sheep's wool is called a '' shearer''. Typically each adult sheep is shorn once each year (a sheep may be said to have been "shorn" or ...
every three years and has to be caught from the wild. When knitted together, the product of the vicuña's wool is very soft and warm. The
Inca The Inca Empire (also known as the Incan Empire and the Inka Empire), called ''Tawantinsuyu'' by its subjects, ( Quechua for the "Realm of the Four Parts",  "four parts together" ) was the largest empire in pre-Columbian America. The adm ...
valued vicuñas highly for their wool, and it was against the law for anyone but royalty to wear vicuña garments; today, the vicuña is the
national animal This is a list of countries that have officially designated one or more animals as their national animals. National animal {, class="wikitable sortable" ! Country ! Name of animal ! Scientific name (Latin name) ! class="unsortable", Picture ...
of
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
and appears on the Peruvian coat of arms. Both under the rule of the Inca and today, vicuñas have been protected by law, but they were heavily hunted in the intervening period. At the time they were declared
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
in 1974, only about 6,000 animals were left. Today, the vicuña population has recovered to about 350,000, and although conservation organizations have reduced its level of threat classification, they still call for active conservation programs to protect populations from poaching,
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
, and other threats. Previously the vicuña was thought not to have been domesticated, and the llama and the alpaca were both regarded as descendants of the closely related guanaco. But DNA research published in 2001 has shown the alpaca may well have vicuña parentage. Today, the vicuña is mainly wild, but the local people still perform special rituals with these creatures, including a fertility rite.


Description

The vicuña is considered more delicate and gracile than the
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco ...
, and smaller. A key distinguishing element of morphology is the better-developed incisor roots for the guanaco. The vicuña's long, woolly coat is tawny brown on the back, whereas the hair on the throat and chest is white and quite long. The head is slightly shorter than the guanaco's and the ears are slightly longer. The length of head and body ranges from 1.45 to 1.60 m (about 5 ft); shoulder height is from 75 to 85 cm (around 3 ft); its weight is from 35 to 65 kg (under 150 lb). It falls prey to the puma and
culpeo The culpeo (''Lycalopex culpaeus''), also known as culpeo zorro, Andean zorro, Andean fox, Paramo wolf, Andean wolf,Comparative ecology of two South American foxes, 'Dusicvon ariseus' and 'culpaeus' by Warren E. Johnson. Doctoral dissertation. Io ...
.


Taxonomy and evolution

There are two subspecies of vicuña: * ''Lama vicugna vicugna'' * ''Lama vicugna mensalis'' While vicuñas are restricted to the more extreme elevations of the Andes in modern times, they may have also been present in the lowland regions of
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
as much as 3500 km south of their current range during the
Late Pleistocene The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch withi ...
and Early Holocene. Fossils of these lowland camelids have been assigned to a species known as ''Lama gracilis'', but genetic and morphological analysis between them and modern vicuña indicate the two may be the same.


Distribution and habitat

Vicuñas are native to the central
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
in South America. They are found in
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
, northwestern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
, and northern
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
. A smaller, introduced population lives in central
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
. Vicuñas live at altitudes of . They feed in daytime on the grassy plains of the Andes Mountains, but spend the nights on the slopes. In these areas, only nutrient-poor, tough, bunch grasses and ''
Festuca ''Festuca'' (fescue) is a genus of flowering plants belonging to the grass family Poaceae (subfamily Pooideae). They are evergreen or herbaceous perennial tufted grasses with a height range of and a cosmopolitan distribution, occurring on ever ...
'' grow. The sun's rays are able to penetrate the thin
atmosphere An atmosphere () is a layer of gas or layers of gases that envelop a planet, and is held in place by the gravity of the planetary body. A planet retains an atmosphere when the gravity is great and the temperature of the atmosphere is low. A ...
, producing relatively warm temperatures during the day; however, the temperatures drop to freezing at night. The vicuña's thick but soft coat is a special adaptation which traps layers of warm air close to its body, so it can tolerate freezing temperatures. Chief predators include pumas and the
culpeo The culpeo (''Lycalopex culpaeus''), also known as culpeo zorro, Andean zorro, Andean fox, Paramo wolf, Andean wolf,Comparative ecology of two South American foxes, 'Dusicvon ariseus' and 'culpaeus' by Warren E. Johnson. Doctoral dissertation. Io ...
.


Behavior

The behavior of vicuñas is similar to that of the guanacos. They are very shy animals, and are easily aroused by intruders, due, among other things, to their extraordinary hearing. Like the guanacos, they frequently lick
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an ad ...
stones and rocks, which are rich in salt, and also drink salt water. Their diets consist mainly of low grasses which grow in clumps on the ground. Vicuñas live in family-based groups made up of a male, five to 15 females, and their young. Each group has its own territory of about 18 km2, which can fluctuate depending on the availability of food.
Mating In biology, mating is the pairing of either opposite- sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reprod ...
usually occurs in March–April, and after a
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pr ...
period of about 11 months, the female gives birth to a single fawn, which is nursed for about 10 months. The fawn becomes independent at about 12 to 18 months old. Young males form bachelor groups and the young females search for a sorority to join. This deters intraspecific competition and inbreeding.


Conservation

From the period of Spanish conquest to 1964, hunting of the vicuña was unrestricted, which reduced its numbers to only 6,000 in the 1960s. As a result, the species was declared endangered in 1974, and its status prohibited the trade of vicuña wool. In Peru, during 1964–1966, the Servicio Forestal y de Caza in cooperation with the US Peace Corps, Nature Conservancy, World Wildlife Fund, and the
National Agrarian University The National Agrarian University, also formally called National Agrarian University – La Molina (Spanish: ''Universidad Nacional Agraria La Molina'') (UNALM), is a public university in Lima, Peru. It is the country's best university in the f ...
of La Molina established a nature conservatory for the vicuña called the Pampa Galeras – Barbara D'Achille in
Lucanas Province Lucanas is the largest province in the Ayacucho Region in Peru. Its seat is Puquio. Geography Some of the highest mountains of the province are Rasuwillka and Qarwarasu. Other mountains are listed below: Some of the largest lakes of the pr ...
,
Ayacucho Ayacucho (, qu, Ayak'uchu) is the capital city of Ayacucho Region and of Huamanga Province, Ayacucho Region, Peru. During the Inca Empire and Viceroyalty of Peru periods the city was known by the name of Huamanga (Quechua: Wamanga), and it co ...
. During that time, a game warden academy was held in Nazca, where eight men from Peru and six from Bolivia were trained to protect the vicuña from poaching. To cooperate on the conservation of the vicuña, the governments of Bolivia and Peru signed the Convention for the Conservation of the Vicuña on 16 August 1969 in
La Paz La Paz (), officially known as Nuestra Señora de La Paz (Spanish pronunciation: ), is the seat of government of the Plurinational State of Bolivia. With an estimated 816,044 residents as of 2020, La Paz is the third-most populous city in Bol ...
, explicitly leaving the treaty open to accession by Argentina and Chile. Convenio para la Conservación de la Vicuña (Wikisource). Source: Registro Oficial N° 144 de 4 de agosto de 1976 (Official Gazette of the Government of Ecuador no. #144, 4 August 1976). Ecuador acceded on 11 February 1976. The Convention prohibited international trade in the vicuña, domestic exploitation of the vicuña, and ordered the parties to create reserves and breeding centres. A follow-up treaty, the Convention for the Conservation and Management of the Vicuña, was signed between Bolivia, Chile, Ecuador and Peru on 20 December 1979 in
Lima Lima ( ; ), originally founded as Ciudad de Los Reyes (City of The Kings) is the capital and the largest city of Peru. It is located in the valleys of the Chillón, Rímac and Lurín Rivers, in the desert zone of the central coastal part of ...
. It explicitly allowed only Argentina to sign it if it also signed the 1969 La Paz Convention (Article 12; Argentina joined in 1981), and did not allow other countries to accede to the convention 'due to its specific character' (Article 13). The 1979 Convention did allow the use of the vicuña under strict circumstances if the animal population had recovered sufficiently. In combination with
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
(effective in 1975), as well as USA and EU trade legislation, the Conventions were extremely successful, as the vicuña population substantially grew as a result. The estimated population in Peru increased from 6,000 to 75,000 with protection by game wardens. Currently, the community of Lucanas conducts a ''chaccu'' (herding, capturing, and shearing) on the reserve each year to harvest the wool, organized by the National Council for South American Camelids (CONACS). In Bolivia, the Ulla Ulla National Reserve was founded in 1977 partly as a sanctuary for the species. Their numbers grew to 125,000 in Peru, Chile, Argentina, and Bolivia. Since this was a ready "cash crop" for community members, the countries relaxed regulations on vicuña wool in 1993, enabling its trade once again. The wool is sold on the world market for over $300 per kg, to help support the community. In 2002, the
US Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
reclassified most populations as threatened, but still lists Ecuador's population as endangered. While the population levels have recovered to a healthy level, poaching remains a constant threat, as do habitat loss and other threats. Consequently, the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
still supports active conservation programs to protect vicuñas, though they lowered their status to least concern in 2018.


Vicuña wool

The wool is popular due to its warmth, and is used for apparel such as socks, sweaters, accessories, shawls, coats, and suits, and home furnishings such as blankets and throws. Its properties come from the tiny scales on the hollow, air-filled fibres. It causes them to interlock and trap insulating air. Vicuñas have some of the finest fibers in the world, at a diameter of 12 μm. The fiber of cashmere goats is 14 to 19 μm, while
angora rabbit The Angora rabbit ( tr, Ankara tavşanı), which is one of the oldest types of domestic rabbit, is bred for the long fibers of its coat, known as ''Angora wool'', which are gathered by shearing, combing or plucking. Because rabbits do not posses ...
is 8 to 12 μm and that of shahtoosh from the Tibetan antelope, or chiru, is from 9 to 12 μm.


Gallery

File:Escudo nacional del Perú.svg, Vicuña in the coat of arms of Peru File:Alpaca, Llama, Vicuna (illustration from New Student's Reference Work, 1914).jpg, Comparison of alpaca, llama, and vicuña (1914) File:Vicuña.jpg, A vicuña on rocky terrain (2008) File:Vicuñas camino a Geysers El Tatio.jpg, Vicuñas on road to
El Tatio El Tatio is a geothermal field with many geysers located in the Andes Mountains of northern Chile at above mean sea level. It is the third-largest geyser field in the world and the largest in the Southern Hemisphere. Various meanings have been ...
geysers,
San Pedro de Atacama San Pedro de Atacama is a Chilean town and commune in El Loa Province, Antofagasta Region. It is located east of Antofagasta, some 106 km (60 mi) southeast of Calama and the Chuquicamata copper mine, overlooking the Licancabur volcano. ...
,
Antofagasta Region The Antofagasta Region ( es, Región de Antofagasta, ) is one of Chile's sixteen first-order administrative divisions. The second-largest region of Chile in area, it comprises three provinces, Antofagasta, El Loa and Tocopilla. It is bordered t ...
,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
(2011) File:Female Vicuña running.jpg, Vicuña in
Jujuy Province Jujuy is a province of Argentina, located in the extreme northwest of the country, at the borders with Chile and Bolivia. The only neighboring Argentine province is Salta to the east and south. Geography There are three main areas in Jujuy: * ...
in the Argentine Altiplano (2011) File:Vicugna-ESO.jpg, Vicuña fawn in the
Atacama Desert The Atacama Desert ( es, Desierto de Atacama) is a desert plateau in South America covering a 1,600 km (990 mi) strip of land on the Pacific coast, west of the Andes Mountains. The Atacama Desert is the driest nonpolar desert in th ...
(2014) File:Vicugna vicugna Salar de Chalviri.jpg, Vicunas in Salar de Chalviri, Bolivia


References


Notes


External links

* * * *
Convention for the Conservation of the Vicuña (La Paz 1969, including the 1976 Ecuadorian accession)
– Spanish Wikisource
Convention for the Conservation and Management of the Vicuña (Lima 1979)
– Spanish Wikisource
Convention for the Conservation and Management of the Vicuña (Lima 1979)
– Original scan a
Conveniovicuna.org
{{DEFAULTSORT:Vicuna Camelids Mammals of the Andes Mammals of Argentina Mammals of Bolivia Mammals of Chile Mammals of Peru Wool animals Andean studies Mammals described in 1782 Taxa named by Juan Ignacio Molina Taxobox binomials not recognized by IUCN National symbols of Peru