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Tusks are elongated, continuously growing front
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, ...
that protrude well beyond the
mouth In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on ...
of certain
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
species. They are most commonly canine teeth, as with pigs and
walrus The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fami ...
es, or, in the case of
elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantida ...
s, elongated
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s. Tusks share common features such as extra-oral position, growth pattern, composition and structure, and lack of contribution to ingestion. Tusks are thought to have adapted to the extra-oral environments, like dry or aquatic or arctic. In most tusked species both the males and the females have tusks although the males' are larger. Most mammals with tusks have a pair of them growing out from either side of the mouth. Tusks are generally curved and have a smooth, continuous surface. The male
narwhal The narwhal, also known as a narwhale (''Monodon monoceros''), is a medium-sized toothed whale that possesses a large " tusk" from a protruding canine tooth. It lives year-round in the Arctic waters around Greenland, Canada and Russia. It is ...
's straight single helical tusk, which usually grows out from the left of the mouth, is an exception to the typical features of tusks described above. Continuous growth of tusks is enabled by formative tissues in the apical openings of the roots of the teeth. Prior to over hunting and proliferation of the ivory trade, elephant tusks weighing over were not uncommon, though it is rare today to see any over . Other than mammals,
dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typic ...
s are the only known vertebrates to have true tusks.


Function

Tusks have a variety of uses depending on the animal. Social displays of dominance, particularly among males, are common, as is their use in defense against attackers. Elephants use their tusks as digging and boring tools. Walruses use their tusks to grip and haul out on ice. It has been suggested that tusk's structure has evolved to be compatible with extra-oral environments.


Use by humans

Tusks are used by humans to produce
ivory Ivory is a hard, white material from the tusks (traditionally from elephants) and teeth of animals, that consists mainly of dentine, one of the physical structures of teeth and tusks. The chemical structure of the teeth and tusks of mammals i ...
, which is used in artifacts and
jewellery Jewellery ( UK) or jewelry ( U.S.) consists of decorative items worn for personal adornment, such as brooches, rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the clothes. From a w ...
, and formerly in other items such as
piano The piano is a stringed keyboard instrument in which the strings are struck by wooden hammers that are coated with a softer material (modern hammers are covered with dense wool felt; some early pianos used leather). It is played using a keyboa ...
keys. Consequently, many tusk-bearing species have been hunted commercially and several are
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
. The
ivory trade The ivory trade is the commercial, often illegal trade in the ivory tusks of the hippopotamus, walrus, narwhal, mammoth, and most commonly, African and Asian elephants. Ivory has been traded for hundreds of years by people in Africa and Asia ...
has been severely restricted by the
United Nations The United Nations (UN) is an intergovernmental organization whose stated purposes are to maintain international peace and security, develop friendly relations among nations, achieve international cooperation, and be a centre for harmoni ...
Convention on International Trade in Endangered Species of Wild Fauna and Flora CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
(CITES). Tusked animals in human care may undergo tusk trimming or removal for health and safety concerns. Furthermore, surgical veterinary procedures to remove tusks have been explored to mitigate human-wildlife conflicts.


Gallery


See also

*
Fang A fang is a long, pointed tooth. In mammals, a fang is a modified maxillary tooth, used for biting and tearing flesh. In snakes, it is a specialized tooth that is associated with a venom gland (see snake venom). Spiders also have external fa ...
, a long canine tooth (in mammals) *
Ivory trade The ivory trade is the commercial, often illegal trade in the ivory tusks of the hippopotamus, walrus, narwhal, mammoth, and most commonly, African and Asian elephants. Ivory has been traded for hundreds of years by people in Africa and Asia ...
*
Eco-economic decoupling In economic and environmental fields, decoupling refers to an economy that would be able to grow without corresponding increases in environmental pressure. In many economies, increasing production ( GDP) currently raises pressure on the environ ...


References

{{Reflist Mammal anatomy Bone products Ivory