Stromatoporoidea is an extinct clade of

sea sponges Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through ...

common in the fossil record from the

Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...

through the Devonian. They were especially abundant and important reef-formers in the Silurian and most of the Devonian.Stock, C.W. 2001, Stromatoporoidea, 1926–2000: ''Journal of Paleontology'', v. 75, p. 1079–1089. The group was previously thought to be related to the

coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and ...

s and placed in the phylum

Cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that ...

. They are now classified in the phylum Porifera, specifically the

sclerosponge Demosponges (Demospongiae) are the most diverse class in the phylum Porifera. They include 76.2% of all species of sponges with nearly 8,800 species worldwide (World Porifera Database). They are sponges with a soft body that covers a ha ...

s. There are numerous

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

forms with spherical, branching or encrusting skeletons of laminated calcite with vertical ''pillars'' between the ''laminae''. Specimen of its oldest genus, ''Priscastroma'', have been found within the Middle Ordovician Sediments. This same genus has been referred to as the species ''P. gemina'' Khrom., and is known to have been known to branch off into two forms, A and B. Form A gave rise to the genus ''Cystostroma'' while form B gave rise to the genus ''Labechia'' and its descendants. Paleozoic stromatoporoids died out at the

Hangenberg Event The Hangenberg event, also known as the Hangenberg crisis or end-Devonian extinction, is a mass extinction that occurred at the end of the Famennian stage, the last stage in the Devonian Period (roughly 358.9 ± 0.4 million years ago). It is usuall ...

at the end of the Devonian. Purported

Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...

stromatoporoids may be unrelated, thus making "stromatoporoids" a

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...

group if they are included. Stromatoporoids are useful markers whose form and occurrence can diagnose the

depositional environment In geology, depositional environment or sedimentary environment describes the combination of physical, chemical, and biological processes associated with the deposition of a particular type of sediment and, therefore, the rock types that will be ...

of sedimentary strata. Paleozoic stromatoporoids hosted various symbiotic endobionts such as

rugosa The rugosa, also called the tetracorallia or horn coral, are an extinct order of solitary and colonial corals that were abundant in Middle Ordovician to Late Permian seas. Solitary rugosans (e.g., '' Caninia'', '' Lophophyllidium'', '' Neoza ...

ns, syringoporids, worms and

cornulitid Cornulitida is an extinct order of encrusting animals from class Tentaculita, which were common around the globe in the Ordovician to Devonian oceans, and survived until the Carboniferous. The organisms had shells, and were subject to predation ...

s. Image:StromatoporoidSideDevColumbus.jpg, Side view of a stromatoporoid showing ''laminae'' and ''pillars'';

Columbus Limestone The Columbus Limestone is a mapped bedrock unit consisting primarily of fossiliferous limestone, and it occurs in Ohio, Pennsylvania, and Virginia in the United States, and in Ontario, Canada. Description Depositional environment The depositiona ...

( Devonian) of Ohio. File:Stromatoporoid1 Keyser Formation.jpg, Stromatoporoids in the Devonian

Keyser Formation The Late Silurian to Early Devonian Keyser Formation is a mapped limestone bedrock unit in Pennsylvania, Maryland, Virginia, and West Virginia. Description The Keyser is a nodular limestone overlain by thick- and thin-bedded limestone and lam ...

, Pennsylvania. File:Densastroma pexisum.jpg, ''Densastroma pexisum'', a stromatoporoid from the Silurian of

Saaremaa Island Saaremaa is the largest island in Estonia, measuring . The main island of Saare County, it is located in the Baltic Sea, south of Hiiumaa island and west of Muhu island, and belongs to the West Estonian Archipelago. The capital of the islan ...

Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...

. File:Stromatoporoid reef, Cairn Formation (Devonian).jpg, Stromatoporoid reef (

Cairn Formation The Cairn Formation is a geologic formation of Late Devonian (Frasnian) age in the Western Canada Sedimentary Basin. It was named for the Cairn River near its junction with the Southesk River in Jasper National Park by D.J. McLaren in 1955.McLare ...

, Late Devonian) in

Calgary, Alberta Calgary ( ) is the largest city in the western Canadian province of Alberta and the largest metro area of the three Prairie Provinces. As of 2021, the city proper had a population of 1,306,784 and a metropolitan population of 1,481,806, makin ...

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

References

External links

University of California, Berkeley 2019 museum blog post on StromatoporoideaDigital Atlas of Ancient Life page on Stromatoporoidea
Ordovician invertebrates Devonian animals Late Devonian animals Silurian animals Prehistoric animal classes {{paleo-sponge-stub