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A stinger (or sting) is a sharp
organ Organ may refer to: Biology * Organ (biology), a part of an organism Musical instruments * Organ (music), a family of keyboard musical instruments characterized by sustained tone ** Electronic organ, an electronic keyboard instrument ** Hammond ...
found in various
animal Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage ...
s (typically
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s and other
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s) capable of injecting
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a st ...
, usually by piercing the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water rel ...
of another animal. An
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
sting is complicated by its introduction of
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a st ...
, although not all stings are venomous. Bites, which can introduce
saliva Saliva (commonly referred to as spit) is an extracellular fluid produced and secreted by salivary glands in the mouth. In humans, saliva is around 99% water, plus electrolytes, mucus, white blood cells, epithelial cells (from which DNA can ...
as well as additional pathogens and diseases, are often confused with stings, and vice versa. Specific components of venom are believed to give rise to an
allergic reaction Allergies, also known as allergic diseases, refer a number of conditions caused by the hypersensitivity of the immune system to typically harmless substances in the environment. These diseases include hay fever, food allergies, atopic derm ...
, which in turn produces skin lesions that may vary from a small itching weal, or slightly elevated area of the skin, to large areas of inflamed skin covered by
vesicles Vesicle may refer to: ; In cellular biology or chemistry * Vesicle (biology and chemistry), a supramolecular assembly of lipid molecules, like a cell membrane * Synaptic vesicle ; In human embryology * Vesicle (embryology), bulge-like features o ...
and crusted
lesions A lesion is any damage or abnormal change in the tissue of an organism, usually caused by disease or trauma. ''Lesion'' is derived from the Latin "injury". Lesions may occur in plants as well as animals. Types There is no designated classifi ...
. Stinging insects produce a painful swelling of the skin, the severity of the lesion varying according to the location of the sting, the identity of the insect and the sensitivity of the subject. Many species of
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfami ...
s and
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
s have two poison glands, one gland secreting a
toxin A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849 ...
in which
formic acid Formic acid (), systematically named methanoic acid, is the simplest carboxylic acid, and has the chemical formula HCOOH and structure . It is an important intermediate in chemical synthesis and occurs naturally, most notably in some ants. Est ...
is one recognized constituent, and the other secreting an
alkaline In chemistry, an alkali (; from ar, القلوي, al-qaly, lit=ashes of the saltwort) is a basic, ionic salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as a base that dissolves in water. A solution of a ...
neurotoxin Neurotoxins are toxins that are destructive to nerve tissue (causing neurotoxicity). Neurotoxins are an extensive class of exogenous chemical neurological insultsSpencer 2000 that can adversely affect function in both developing and mature nerv ...
; acting independently, each toxin is rather mild, but when they combine through the sting, the combination has strong irritating properties. In a small number of cases, the second occasion of a bee or wasp sting causes a severe allergic reaction known as
anaphylaxis Anaphylaxis is a serious, potentially fatal allergic reaction and medical emergency that is rapid in onset and requires immediate medical attention regardless of use of emergency medication on site. It typically causes more than one of the follo ...
. While the overwhelming majority of insects withdraw their stingers from their victims, a few insects leave them in the wounds. For example, of the 20,000 species of bees worldwide, only the half-dozen species of honeybees ('' Apis'') are reported to have a barbed stinger that cannot be withdrawn; of wasps, nearly all are reported to have smooth stingers with the exception of two species, ''
Polybia rejecta ''Polybia rejecta'' is a species of social wasp found in the Neotropics region of the world. It was discovered by Fabricius in South America in the 1790s. The wasp is associated with many other organisms, particularly specific species of ants ...
'' and '' Synoeca surinama''. A few non-insect arthropods, such as
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always en ...
s, also sting.


Arthropods

Among
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s, a sting or stinger is a sharp organ, often connected with a venom gland and adapted to inflict a wound by piercing, as with the caudal sting of a
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always en ...
. Stings are usually located at the rear of the animal. Animals with stings include bees, wasps (including
hornet Hornets (insects in the genus ''Vespa'') are the largest of the eusocial wasps, and are similar in appearance to their close relatives yellowjackets. Some species can reach up to in length. They are distinguished from other vespine wasps by t ...
s) and
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always en ...
s, as well as a single beetle species (''
Onychocerus albitarsis ''Onychocerus albitarsis'' is a relatively rare species of beetle in the family Cerambycidae from the Amazon and Atlantic forest regions in Brazil, Bolivia, Paraguay and southern Peru. It is remarkable for being the only known beetle that has a ...
'') that can deliver a venomous sting from its antennae, whose terminal segments have evolved to resemble a scorpion's tail. In all stinging
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typic ...
the sting is a modified
ovipositor The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
. Unlike most other stings,
honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosm ...
workers' stings are strongly barbed and lodge in the flesh of
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s upon use, tearing free from the honey bee's body, killing the bee within minutes. The sting has its own
ganglion A ganglion is a group of neuron cell bodies in the peripheral nervous system. In the somatic nervous system this includes dorsal root ganglia and trigeminal ganglia among a few others. In the autonomic nervous system there are both sympathe ...
, and it continues to saw into the target's flesh and release venom for several minutes. This trait is of obvious disadvantage to the individual but protects the hive from attacks by large animals; aside from the effects of the venom, the remnant also marks the stung animal with honey bee alarm pheromone. The barbs of a honey bee's attack are only suicidal if the skin is elastic, as is characteristic of vertebrates such as birds and mammals; honey bees can sting other insects repeatedly without dying. The sting of nearly all other bees and other sting-bearing organisms is not barbed and can be used to sting repeatedly. The description of barbed or unbarbed is not precise: there are barbs on the stings of yellowjacket wasps and the Mexican honey wasp, but the barbs are so small that the wasp can sometimes withdraw its sting apparatus from victim's skin. The stings of some wasps, such as those of the '' Polistes versicolor'', contain relatively large amounts of 5-hydroxytryptamine (5-HT) in its venoms. The 5-HT in these venoms has been found to play at least two roles: one as a pain-producing agent and the other in the distribution and penetration of the paralyzing components to vulnerable sites in the offender. This helps in the rapid immobilization of the animal or of the body parts receiving the venom. Spiders only bite, although some tarantulas have barbed bristles called
urticating hairs Urticating hairs or urticating bristles are one of the primary defense mechanisms used by numerous plants, almost all New World tarantulas, and various lepidopteran caterpillars. ''Urtica'' is Latin for "nettle" (stinging nettles are in the genus ...
. Certain caterpillars also have urticating hairs.
Centipede Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...
s also possess a venomous bite rather than a sting, inflicted with a highly modified first pair of legs, called forcipules.


Other animals

Organs that perform similar functions in non-arthropods are often referred to as "stings". These organs include the modified
dermal denticle A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as ...
of the
stingray Stingrays are a group of sea rays, which are cartilaginous fish related to sharks. They are classified in the suborder Myliobatoidei of the order Myliobatiformes and consist of eight families: Hexatrygonidae (sixgill stingray), Plesiobatidae ...
, the venomous spurs on the hind legs of the male
platypus The platypus (''Ornithorhynchus anatinus''), sometimes referred to as the duck-billed platypus, is a semiaquatic, egg-laying mammal endemic to eastern Australia, including Tasmania. The platypus is the sole living representative or mono ...
, and the cnidocyte
tentacle In zoology, a tentacle is a flexible, mobile, and elongated organ present in some species of animals, most of them invertebrates. In animal anatomy, tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work main ...
s of the
jellyfish Jellyfish and sea jellies are the informal common names given to the medusa-phase of certain gelatinous members of the subphylum Medusozoa, a major part of the phylum Cnidaria. Jellyfish are mainly free-swimming marine animals with umbre ...
. The term ''sting'' was historically often used for the
fang A fang is a long, pointed tooth. In mammals, a fang is a modified maxillary tooth, used for biting and tearing flesh. In snakes, it is a specialized tooth that is associated with a venom gland (see snake venom). Spiders also have external fa ...
of a
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
,Oxford English Dictionary, 2nd ed. "Sting ... Applied also to the fang or venom-tooth (and erroneously to the forked tongue) of a poisonous serpent." although this usage is uncommon today. Snakes are said, correctly, to bite, not sting.


See also

* Bee sting * Cnidocyte * Chelicerae * Forcipules * Insect bite * Schmidt Sting Pain Index *
Starr sting pain scale The Starr sting pain scale was created by the entomologist Christopher Starr as a scale to compare the overall pain of hymenopteran stings on a four-point scale, an expansion of the "pain index" originally created by Justin Schmidt. 1 is the lowe ...
* Stinging plant


References


External links

*{{Commons category-inline, Stingers Animal anatomy