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Spatiotemporal patterns are
pattern A pattern is a regularity in the world, in human-made design, or in abstract ideas. As such, the elements of a pattern repeat in a predictable manner. A geometric pattern is a kind of pattern formed of geometric shapes and typically repeated l ...
s that occur in a wide range of natural phenoma and are characterized by a spatial and a temporal patterning. The general rules of
pattern formation The science of pattern formation deals with the visible, ( statistically) orderly outcomes of self-organization and the common principles behind similar patterns in nature. In developmental biology, pattern formation refers to the generation of ...
hold. In contrast to "static", pure spatial patterns, the full complexity of spatiotemporal patterns can only be recognized over time. Any kind of traveling wave is a good example of a spatiotemporal pattern. Besides the shape and
amplitude The amplitude of a periodic variable is a measure of its change in a single period (such as time or spatial period). The amplitude of a non-periodic signal is its magnitude compared with a reference value. There are various definitions of am ...
of the wave (spatial part), its time-varying position (and possibly shape) in space is an essential part of the entire pattern. The distinction between spatial and spatio-temporal patterns in
nature Nature, in the broadest sense, is the physical world or universe. "Nature" can refer to the phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. Although humans ar ...
is not clear-cut because a static, invariable pattern will never occur in the strict sense. Even
rock formation A rock formation is an isolated, scenic, or spectacular surface rock outcrop. Rock formations are usually the result of weathering and erosion sculpting the existing rock. The term ''rock formation'' can also refer to specific sedimen ...
s will slowly change on a time-scale of 10s of millions of years, therefore the distinction lies in the time scale of change in relation to human
experience Experience refers to conscious events in general, more specifically to perceptions, or to the practical knowledge and familiarity that is produced by these conscious processes. Understood as a conscious event in the widest sense, experience invol ...
. Already the snapshot state of a
dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...
will usually be taken as an example of a purely spatial pattern although this is clearly not the case. It is thus apt to say that spatiotemporal patterns in nature are the rule rather than the exception.


Physics

Many hydrodynamical systems show s.t. pattern formation: *
Rayleigh–Bénard convection In fluid thermodynamics, Rayleigh–Bénard convection is a type of natural convection, occurring in a planar horizontal layer of fluid heated from below, in which the fluid develops a regular pattern of convection cells known as Bénard cells. ...
*
Taylor–Couette flow In fluid dynamics, the Taylor–Couette flow consists of a viscous fluid confined in the gap between two rotating cylinders. For low angular velocities, measured by the Reynolds number ''Re'', the flow is steady and purely azimuthal. This basic ...
*
Liquid crystal Liquid crystal (LC) is a state of matter whose properties are between those of conventional liquids and those of solid crystals. For example, a liquid crystal may flow like a liquid, but its molecules may be oriented in a crystal-like way. T ...
instabilities


Chemistry

Any type of reaction–diffusion system that produces spatial patterns will also, due to the time-dependency of both reactions and diffusion, produce spatiotemporal patterns.


Biology


Neurobiology

Neural networks, both
artificial Artificiality (the state of being artificial or manmade) is the state of being the product of intentional human manufacture, rather than occurring naturally through processes not involving or requiring human activity. Connotations Artificiality ...
and
natural Nature, in the broadest sense, is the physics, physical world or universe. "Nature" can refer to the phenomenon, phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. ...
, produce a virtually unbounded variety of s.t. patterns, both in
sensory perception Perception () is the organization, identification, and interpretation of sensory information in order to represent and understand the presented information or environment. All perception involves signals that go through the nervous system ...
, learning, thinking and reasoning as well as in spontaneous activity. It has for example been demonstrated that spiral waves, signatures of many excitable systems can occur in neocortical preparations.


Communication

All
communication Communication (from la, communicare, meaning "to share" or "to be in relation with") is usually defined as the transmission of information. The term may also refer to the message communicated through such transmissions or the field of inqu ...
,
language Language is a structured system of communication. The structure of a language is its grammar and the free components are its vocabulary. Languages are the primary means by which humans communicate, and may be conveyed through a variety of ...
, relies on spatiotemporal
encoding In communications and information processing, code is a system of rules to convert information—such as a letter, word, sound, image, or gesture—into another form, sometimes shortened or secret, for communication through a communicatio ...
of information, producing and
transmitting In electronics and telecommunications, a radio transmitter or just transmitter is an electronic device which produces radio waves with an antenna. The transmitter itself generates a radio frequency alternating current, which is applied to th ...
sound In physics, sound is a vibration that propagates as an acoustic wave, through a transmission medium such as a gas, liquid or solid. In human physiology and psychology, sound is the ''reception'' of such waves and their ''perception'' by ...
variations or any type of
signal In signal processing, a signal is a function that conveys information about a phenomenon. Any quantity that can vary over space or time can be used as a signal to share messages between observers. The '' IEEE Transactions on Signal Processing' ...
i.e. single building blocks of information that are varied over time. -Even though
written language A written language is the representation of a spoken or gestural language by means of a writing system. Written language is an invention in that it must be taught to children, who will pick up spoken language or sign language by exposure eve ...
appears to exist only as a (2D) spatial
concatenation In formal language theory and computer programming, string concatenation is the operation of joining character strings end-to-end. For example, the concatenation of "snow" and "ball" is "snowball". In certain formalisations of concatena ...
of letters -
string String or strings may refer to: * String (structure), a long flexible structure made from threads twisted together, which is used to tie, bind, or hang other objects Arts, entertainment, and media Films * ''Strings'' (1991 film), a Canadian ani ...
s, it must be decoded sequentially over time. Any kind of language that is understood by organisms is thus eventually a transcoding of neural s.t. signals and will - in successful communication - evoke similar patterns of neural activity in the recipient as they existed in the sender. For example, the warning call of a bird when it perceives a predator will produce a similar type and degree of alarmedness (eventually a certain kind of neural activity pattern) in other individuals even though they have not yet seen or heard the potential attacker. Even artificial languages, e.g.
computer language A computer language is a formal language used to communicate with a computer. Types of computer languages include: * Construction language – all forms of communication by which a human can specify an executable problem solution to a compu ...
s, are not read and interpreted in one step, but sequentially, thus, their meaningfully arranged vocabulary (e.g. "
computer code A computer is a machine that can be programmed to carry out sequences of arithmetic or logical operations (computation) automatically. Modern digital electronic computers can perform generic sets of operations known as programs. These progra ...
") can be seen as a s.t. pattern.


Genetics

As a particular type of language, the "static" (neglecting random
transcription error A transcription error is a specific type of data entry error that is commonly made by human operators or by optical character recognition (OCR) programs. Human transcription errors are commonly the result of typographical mistakes; putting one� ...
s, recombination and
mutation In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, ...
) DNA and its
transcription Transcription refers to the process of converting sounds (voice, music etc.) into letters or musical notes, or producing a copy of something in another medium, including: Genetics * Transcription (biology), the copying of DNA into RNA, the fir ...
pattern over time yields biologically essential s.t. patterns.
Gene regulatory network A gene (or genetic) regulatory network (GRN) is a collection of molecular regulators that interact with each other and with other substances in the cell to govern the gene expression levels of mRNA and proteins which, in turn, determine the fun ...
s are responsible for regulation the time course of
gene expression Gene expression is the process by which information from a gene is used in the synthesis of a functional gene product that enables it to produce end products, protein or non-coding RNA, and ultimately affect a phenotype, as the final effect. T ...
level which can be analyzed using
expression profiling In the field of molecular biology, gene expression profiling is the measurement of the activity (the expression) of thousands of genes at once, to create a global picture of cellular function. These profiles can, for example, distinguish between c ...
.


Literature

*


References

{{Reflist Information theory Pattern formation Space and time