Scent gland are

exocrine gland Exocrine glands are glands that secrete substances on to an epithelial surface by way of a duct. Examples of exocrine glands include sweat, salivary, mammary, ceruminous, lacrimal, sebaceous, prostate and mucous. Exocrine glands are one ...

s found in most

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...

s. They produce semi-viscous

secretion 440px Secretion is the movement of material from one point to another, such as a secreted chemical substance from a cell or gland. In contrast, excretion is the removal of certain substances or waste products from a cell or organism. The classic ...

s which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and

sexual behaviour Human sexual activity, human sexual practice or human sexual behaviour is the manner in which humans experience and express their sexuality. People engage in a variety of sexual acts, ranging from activities done alone (e.g., masturbation) t ...

. The odor may be subliminal—not consciously detectable. Though it is not their primary function, the

salivary gland The salivary glands in mammals are exocrine glands that produce saliva through a system of ducts. Humans have three paired major salivary glands ( parotid, submandibular, and sublingual), as well as hundreds of minor salivary glands. Salivary ...

s may also function as scent glands in some animals.

In even-toed ungulates

The

even-toed ungulate The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...

s (Artiodactyla) have many specialized skin glands, the

s of which are involved in semiochemical communication. These glands include the sudoriferous glands (located on the forehead, between the

antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on ...

s and eyes), the

preorbital gland The preorbital gland is a paired exocrine gland found in many species of hoofed animals, which is homologous to the lacrimal gland found in humans. These glands are trenchlike slits of dark blue to black, nearly bare skin extending from the me ...

s (extending from the medial canthus of each eye), the

nasal glands The nasal glands are the seromucous glands in the respiratory The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and pla ...

(located inside the nostrils), the

interdigital gland Scent gland are exocrine glands found in most mammals. They produce semi-viscous secretions which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and ...

s (located between the toes), the

preputial gland Preputial glands are exocrine glands that are located in the folds of skin front of the genitals of some mammals. They occur in several species, including mice, ferrets, rhinoceroses, and even-toed ungulates and produce pheromones. The glands play ...

(located inside the foreskin of the penis), the metatarsal glands (located outside of the hind legs), the tarsal glands (located inside of the hind legs), and the inguinal glands in the lower belly or groin area. Like many other species of Artiodactyla,

deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...

have seven major external scent glands distributed throughout their bodies. Deer rely heavily on these scent glands to communicate with other members of their species, and possibly even with members of other species. For example, male

white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...

(''Odocoileus virginianus'') are often seen working over a scrape. First, the animal scrapes at the dirt with its

hooves The hoof (plural: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits, yet the rumi ...

, depositing the scent from his interdigital gland on the ground. After that, he may bite the tip off an overhanging branch, depositing secretions from his salivary glands onto the branch. He may then rub his face on the overhanging branch, depositing secretions from the sudoriferous and preorbital glands on it. The tarsal gland appears to operate by a different mechanism than the other external scent glands. A behavior called '' rub-urination'' is central to this mechanism. During rub-urination, the animal squats while urinating so that urine will run down the insides of its legs and onto its tarsal glands. The tarsal glands have a tuft of hair which is specially adapted to extract certain chemical compounds from the animal's urine. For example, in the black-tailed deer (''Odocoileus hemionus columbianus''), the major constituent of the tarsal gland secretion is a

lipid Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids in ...

, (Z)-6-dodecen-4-olide. This compound does not originate in the tarsal gland itself, but rather it is extracted from the animal's urine by the tarsal hair tuft during the rub-urination process. In white-tailed deer, the presence and concentration of certain chemical compounds in the urine depend on the season, gender, reproductive status and

social rank A social class is a grouping of people into a set of hierarchical social categories, the most common being the upper, middle and lower classes. Membership in a social class can for example be dependent on education, wealth, occupation, inc ...

of the animals. This fact, along with the observation of rub-urination behavior in this animal (at least in the male) indicates that urine probably plays a role in

olfactory communication Animal communication is the transfer of information from one or a group of animals (sender or senders) to one or more other animals (receiver or receivers) that affects the current or future behavior of the receivers. Information may be sent int ...

in deer.

In carnivorans

Canids have several scent glands that are used in

. The fossa has several scent glands. Like herpestids it has a

perianal The anus ( Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, ...

skin gland inside an anal sac which surrounds the anus like a pocket. The pocket opens to the exterior with a horizontal slit below the tail. Other glands are located near the penis or vagina, with the penile glands emitting a strong odor. Like the herpestids, it has no prescrotal glands.

Anal gland Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving ...

s are found in all

carnivora Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...

including

wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...

bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the No ...

sea otter The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of the weasel family, but among the smal ...

s and kinkajous.

In other animals

Apocrine sweat gland An apocrine sweat gland (; from Greek ''apo'' 'away' and ''krinein'' 'to separate') is composed of a coiled secretory portion located at the junction of the dermis and subcutaneous fat, from which a straight portion inserts and secretes into the ...

s, such as in the

armpits The axilla (also, armpit, underarm or oxter) is the area on the human body directly under the shoulder joint. It includes the axillary space, an anatomical space within the shoulder girdle between the arm and the thoracic cage, bounded superior ...

of humans *

Sebaceous gland A sebaceous gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in the greatest n ...

s, such as the

cranial Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...

surface glands of the red-bellied lemur *

Flank gland The flank gland is one of several specialized posterolateral glands (a group of glands located along the sides and the rear) found predominately in certain species of rodents including voles, shrews, hamsters, and other members of the family Criceti ...

s, such as in

vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...

s or

shrew Shrews (family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to differ ...

s * Various glands producing pheromone signals found in nonavian reptiles, including

Amphisbaenia Amphisbaenia (called amphisbaenians or worm lizards) is a group of usually legless squamates, comprising over 200 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. As ...

ns,

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia altho ...

snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...

s, and Crocodylians * Castor sacs, found in

beaver Beavers are large, semiaquatic rodents in the genus ''Castor'' native to the temperate Northern Hemisphere. There are two extant species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers a ...

s * Perineal glands, found in viverrids, guinea pigs, and porcupines *

Preputial glands Preputial glands are exocrine glands that are located in the folds of skin front of the genitals of some mammals. They occur in several species, including mice, ferrets, rhinoceroses, and even-toed ungulates and produce pheromones. The glands pl ...

, found in many species including mice, pigs, and wolves * Supracaudal glands, found in canidsFox, Michael W., and James A. Cohen.
Canid communication
" How animals communicate (1977): 728-748.

References

{{DEFAULTSORT:Scent Gland Glands Mammal anatomy

In even-toed ungulates

In carnivorans

In other animals

See also

References