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The savannah monitor (''Varanus exanthematicus'') is a medium-sized
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
monitor lizard Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and one species is also found in the Americas as an invasive species. About 80 species are rec ...
native to
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. The species is known as Bosc's monitor in Europe, since French scientist Louis Bosc first described the species. It belongs to the
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between ...
'' Polydaedalus''.


Etymology

The specific name ''exanthematicus'' is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''exanthem'' , meaning an eruption or blister of the skin.
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
and
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...
Louis Augustin Guillaume Bosc Louis Augustin Guillaume Bosc (or Louis-Augustin Bosc d'Antic) (29 January 1759 – 10 July 1828) was a French botanist, invertebrate zoologist, and entomologist. Biography Bosc was born in Paris, the son of Paul Bosc d’Antic, a medical doctor ...
originally described this lizard as ''Lacerta exanthematica'' in reference to the large oval
scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number w ...
on the back of its neck. The species was formerly known as ''Lacerta exanthematicus''.


Description

Savannah monitors are stoutly built, with relatively short limbs and toes, and skulls and dentition adapted to feed on hard-shelled prey. They are robust creatures, with powerful limbs for digging, powerful jaws and blunt, peglike teeth. Maximum size is rarely more than 100 cm. The skin coloration pattern varies according to the local habitat substrate. The body scales are large, usually less than 100 scales around midbody, a partly laterally compressed tail with a double dorsal ridge and nostrils equidistant from the eyes and the tip of the snout. The savannah monitor is often confused with the white-throat monitor ''(Varanus albigularis)'', which can grow to lengths of 5–6 ft. While similar in overall appearance, this species possesses significant morphological and ecological differences and is recognized as a very distinct species.


Behaviour


Diet

Their diet is much more restricted than that of other African monitor lizards, consisting mainly of snails, crabs, scorpions, millipedes, centipedes, orthopterans, mantids, hymenopterans, lepidopterans, beetles and other invertebrates, as well as frogs. Information about the diet of savannah monitors in the wild has been recorded in Senegal and Ghana. It feeds almost exclusively on
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s and molluscs. In Senegal, ''Julus'' millipedes were the most common prey of adults; in Ghana, small crickets formed the bulk of the diet of animals less than 2 months old; orthopterans (especially '' Brachytrupes''), scorpions and amphibians were the most common prey of animals 6–7 months old. Many adults also consume large quantities of snails. Full grown ''V. exanthematicus'' have teeth that are quite blunt to help them crack and eat snails. The jaw has evolved to put maximum leverage at the back of the jaw to crush snail shells. Adults will also eat carrion if they come across it. Wild savannah monitors are also known to occasionally eat lizard eggs (such as those of
agamid Agamidae is a family of over 300 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards. Overview Phylogenetically, they may be sister to the ...
s and their own kind).


Reproduction

Females dig a deep hole in the substrate, in which up to 40 or more eggs are laid, which hatch after about 156–160 days. Hatchlings start feeding a few days after the yolk sac has been absorbed, which may take 12 days or more after hatching.


In captivity

The savannah monitor is the most common monitor lizard species available in the pet trade, accounting for almost half (48.0552%) of the entire international trade in live monitor lizards. Despite its prevalence in global pet trade, successful captive reproduction is very rare, and a high mortality rate is associated with the species. Mampam Conservation. Facts and Fiction about Savannah Monitors. Bennett. https://web.archive.org/web/20171214182359/http://www.mampam.com/index.php?option=com_content&task=view&id=172&Itemid=87 Adult specimens frequently become unwanted pets and are reported as being the most common monitor lizards by animal rescue agencies. The skins are traded within the international leather trade and originate mainly from Chad, Mali and Sudan.


Range

Its range extends throughout sub-Saharan Africa from
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
east to
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
and south almost to the Congo River and Rift Valley, where they are replaced by '' V. albigularis''. ''V. exanthematicus'' is primarily a ground-dwelling species that shelters in burrows, although it is sometimes found in bushes or low trees. In the coastal plain of Ghana, ''V. exanthematicus'' juveniles are often associated with the burrows of the giant cricket '' Brachytrupes''.


Threats

''V. exanthematicus'' is listed as least concern by IUCN. The species is hunted for its leather and meat, and for the international pet trade. The trade in wild collected savannah monitors is not of a global conservation concern, due to the vast range of the species; in addition to the collection for the pet trade often occurring over a relatively small area. An average of 30,574 live specimens were imported into the US each year, between 2000 and 2009; total imports of live specimens into the US between 2000 and 2010 was 325,480 animals. During the same period, 1,037 skins, shoes, and products of the species were imported into the US. Trade in live animals comes mainly from
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and Tog ...
(235,903 animals exported between 2000 and 2010),
Togo Togo (), officially the Togolese Republic (french: République togolaise), is a country in West Africa. It is bordered by Ghana to the west, Benin to the east and Burkina Faso to the north. It extends south to the Gulf of Guinea, where its c ...
(188,110 animals exported between 2000 and 2010), and
Benin Benin ( , ; french: Bénin , ff, Benen), officially the Republic of Benin (french: République du Bénin), and formerly Dahomey, is a country in West Africa. It is bordered by Togo to the west, Nigeria to the east, Burkina Faso to the nort ...
(72,964 animals exported between 2000 and 2010). During the same period, total worldwide declared exports of skins and products of the species totalled 37,506. However, substantial undeclared trade in the species occurs from Sudan, Nigeria, and elsewhere.


References


Further reading

* *Bennett, Daniel; Thakoordyal, Ravi (2003). ''The Savannah Monitor Lizard: the truth about ''Varanus exanthematicus. Glossop, England: Viper Press. 84 pp. . * (1993). "The Savanna Monitor (''Varanus exanthematicus'') in Africa and in your home". The Iowa Herpetological Society, June: 2-4 (Reprinted in International Reptile Breeders Association (IRBA), ''Monitor'' 1 (2): 10–12, 1994).


External links


Photo of a Savannah monitor.
* Savmon.or

Modern captive and known ecological information about ''Varanus exanthematicus''. {{DEFAULTSORT:Savannah monitor Varanus Lizards of Africa Reptiles of Uganda Reptiles described in 1792 Taxa named by Louis Augustin Guillaume Bosc Reptiles as pets