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A ruderal species is a
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae excl ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
that is first to colonize disturbed lands. The disturbance may be natural for example,
wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identi ...
s or
avalanche An avalanche is a rapid flow of snow down a slope, such as a hill or mountain. Avalanches can be set off spontaneously, by such factors as increased precipitation or snowpack weakening, or by external means such as humans, animals, and ea ...
sor the consequences of human activities, such as construction ( of roads, of buildings,
mining Mining is the extraction of valuable minerals or other geological materials from the Earth, usually from an ore body, lode, vein, seam, reef, or placer deposit. The exploitation of these deposits for raw material is based on the econom ...
, etc.) or agriculture (abandoned fields,
irrigation Irrigation (also referred to as watering) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has been devel ...
, etc.). The term '' ruderal'' originates from the Latin word '' rudus'', meaning "
rubble Rubble is broken stone, of irregular size, shape and texture; undressed especially as a filling-in. Rubble naturally found in the soil is known also as 'brash' (compare cornbrash)."Rubble" def. 2., "Brash n. 2. def. 1. ''Oxford English Dictionar ...
". Ruderal species typically dominate the disturbed area for a few years, gradually losing the
competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, ind ...
to other native species. However, in extreme disturbance circumstances, such as when the natural
topsoil Topsoil is the upper layer of soil. It has the highest concentration of organic matter and microorganisms and is where most of the Earth's biological soil activity occurs. Description Topsoil is composed of mineral particles and organic matt ...
is covered with a foreign substance, a single-species ruderal community may become permanently established. In addition, some ruderal
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...
may have such a competitive advantage over the native species that they, too, may permanently prevent a disturbed area from returning to its original state despite natural topsoil.


Features

Features contributing to a species' success as ruderal are: * Massive seed production *
Seedling A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (emb ...
s whose
nutritional Nutrition is the biochemical and physiological process by which an organism uses food to support its life. It provides organisms with nutrients, which can be metabolized to create energy and chemical structures. Failure to obtain sufficient nu ...
requirements are modest * Fast-growing roots * Independence of mycorrhizae *
Polyploidy Polyploidy is a condition in which the biological cell, cells of an organism have more than one pair of (Homologous chromosome, homologous) chromosomes. Most species whose cells have Cell nucleus, nuclei (eukaryotes) are diploid, meaning they ha ...


Quantification

Ecologists have proposed various scales for quantifying ruderality, which can be defined as the "ability to thrive where there is disturbance through partial or total destruction of plant biomass" (Grime, Hodgson & Hunt, 1988). The ruderality scale of Grime presents values that are readily available, and it takes into account disturbance factors as well as other indicators such as the annual or perennial character of the plants.


See also

*
Edge effect In ecology, edge effects are changes in population or community structures that occur at the boundary of two or more habitats. Areas with small habitat fragments exhibit especially pronounced edge effects that may extend throughout the range. ...
* Hemeroby *
Pioneer species Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so ...
*
Restoration ecology Restoration ecology is the scientific study supporting the practice of ecological restoration, which is the practice of renewing and restoring degraded, damaged, or destroyed ecosystems and habitats in the environment by active human interrupt ...
* Supertramp (ecology) * Examples of ruderal species: ** '' Cannabis ruderalis'' (family Cannabaceae) ** '' Conyza bonariensis'' (family
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...
) ** '' Dittrichia viscosa'' (
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...
) ** '' Nicotiana glauca'' (
Solanaceae The Solanaceae , or nightshades, are a family of flowering plants that ranges from annual and perennial herbs to vines, lianas, epiphytes, shrubs, and trees, and includes a number of agricultural crops, medicinal plants, spices, weeds, and or ...
)


References


External links

*
St. John TV. 1987. SOIL DISTURBANCE AND THE MINERAL NUTRITION OF NATIVE PLANTS in Proceedings of the 2nd Native Plant Revegetation Symposium

Chapin. FS. III. 1980. The mineral nutrition of wild plants. Annu. Rev. Ecol. Syst., 11:233-260.
*{{Cite EB1911 , wstitle=Ruderal , volume=23 , page=814 , short=1 Ecology terminology Habitat Botany