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An omnivore () is an
animal Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage ...
that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may o ...
s,
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
, fat, and
fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
, and metabolize the nutrients and energy of the sources absorbed. Often, they have the ability to incorporate food sources such as algae,
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...
, and
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were am ...
into their diet. Omnivores come from diverse backgrounds that often independently evolved sophisticated consumption capabilities. For instance, dogs evolved from primarily carnivorous organisms ( Carnivora) while pigs evolved from primarily herbivorous organisms ( Artiodactyla). Despite this, physical characteristics such as tooth morphology may be reliable indicators of diet in mammals, with such morphological adaptation having been observed in bears. The variety of different animals that are classified as omnivores can be placed into further sub-categories depending on their feeding behaviors. Frugivores include maned wolves and
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genu ...
s; insectivores include swallows and pink fairy armadillos; granivores include large ground finches and mice. All of these animals are omnivores, yet still fall into special niches in terms of feeding behavior and preferred foods. Being omnivores gives these animals more
food security Food security speaks to the availability of food in a country (or geography) and the ability of individuals within that country (geography) to access, afford, and source adequate foodstuffs. According to the United Nations' Committee on World ...
in stressful times or makes possible living in less consistent environments.


Etymology and definitions

The word ''omnivore'' derives from Latin ''omnis'' 'all' and ''vora'', from ''vorare'' 'to eat or devour', having been coined by the French and later adopted by the English in the 1800s. Traditionally the definition for omnivory was entirely behavioral by means of simply "including both animal and vegetable tissue in the diet." In more recent times, with the advent of advanced technological capabilities in fields like
gastroenterology Gastroenterology (from the Greek gastḗr- “belly”, -énteron “intestine”, and -logía "study of") is the branch of medicine focused on the digestive system and its disorders. The digestive system consists of the gastrointestinal tract ...
, biologists have formulated a standardized variation of omnivore used for labeling a species' actual ability to obtain energy and nutrients from materials. This has subsequently conditioned two context specific definitions. * Behavioral: This definition is used to specify if a species or individual is actively consuming both plant and animal materials. ''(e.g. "
vegans Veganism is the practice of abstaining from the use of animal product—particularly in diet—and an associated philosophy that rejects the commodity status of animals. An individual who follows the diet or philosophy is known as a vegan. Di ...
do not participate in the omnivore based diet.")'' In the fields of nutrition, sociology and psychology the terms “omnivore” & “omnivory” is often used to distinguish prototypical highly diverse human diet patterns from restricted diet patterns that exclude major categories of food. * Physiological: This definition is often used in academia to specify species that have the capability to obtain energy and nutrients from both plant and animal matter. ''(e.g. "humans are omnivores due to their capability to obtain energy and nutrients from both plant and animal materials.")'' The taxonomic utility of omnivore's traditional and behavioral definition is limited, since the diet, behavior, and
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ...
of one omnivorous species might be very different from that of another: for instance, an omnivorous pig digging for roots and scavenging for fruit and carrion is taxonomically and ecologically quite distinct from an omnivorous chameleon that eats leaves and insects. The term "omnivory" is also not always comprehensive because it does not deal with mineral foods such as salt licks and the consumption of plant and animal material for medical purposes which would not otherwise be consumed (i.e. zoopharmacognosy) within non-omnivores.


Classification, contradictions and difficulties

Though Carnivora is a
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
for species classification, no such equivalent exists for omnivores, as omnivores are widespread across multiple taxonomic
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
s. The Carnivora order does not include all carnivorous species, and not all species within the Carnivora taxon are carnivorous. (The members of Carnivora are formally referred to as carnivorans.) It is common to find physiological carnivores consuming materials from plants or physiological herbivores consuming material from animals, e.g. felines eating grass and deer eating birds. From a behavioral aspect, this would make them omnivores, but from the physiological standpoint, this may be due to zoopharmacognosy. Physiologically, animals must be able to obtain both energy and nutrients from plant and animal materials to be considered omnivorous. Thus, such animals are still able to be classified as carnivores and herbivores when they are just obtaining nutrients from materials originating from sources that do not seemingly complement their classification. For instance, it is well documented that animals such as giraffes, camels, and cattle will gnaw on bones, preferably dry bones, for particular minerals and nutrients.
Felines The Felinae are a subfamily of the family Felidae. This subfamily comprises the small cats having a bony hyoid, because of which they are able to purr but not roar. Other authors have proposed an alternative definition for this subfamily: a ...
, which are usually regarded as obligate carnivores, occasionally eat grass to regurgitate indigestibles (e.g. hair, bones), aid with hemoglobin production, and as a laxative. Occasionally, it is found that animals historically classified as carnivorous may deliberately eat plant material. For example, in 2013, it was considered that American alligators (''Alligator mississippiensis'') may be physiologically omnivorous once investigations had been conducted on why they occasionally eat fruits. It was suggested that alligators probably ate fruits both accidentally and deliberately. "Life-history omnivores" is a specialized classification given to organisms that change their eating habits during their life cycle. Some species, such as grazing waterfowl like geese, are known to eat mainly animal tissue at one stage of their lives, but plant matter at another.Maclean, Gordon Lindsay (1993). ''Roberts' Birds of Southern Africa''. Publisher: New Holland. . The same is true for many insects, such as beetles in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Meloidae, which begin by eating animal tissue as
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e, but change to eating plant matter after they mature. Likewise, many mosquito species in early life eat plants or assorted detritus, but as they mature, males continue to eat plant matter and nectar whereas the females (such as those of '' Anopheles'', '' Aedes'' and '' Culex'') also eat blood to reproduce effectively.


Omnivorous species


General

Although cases exist of herbivores eating meat and carnivores eating
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae excl ...
matter, the classification "omnivore" refers to the adaptation and main food source of the species in general, so these exceptions do not make either individual animals or the species as a whole omnivorous. For the concept of "omnivore" to be regarded as a scientific classification, some clear set of measurable and relevant criteria would need to be considered to differentiate between an "omnivore" and other categories, e.g.
faunivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
, folivore, and scavenger. Some researchers argue that evolution of any species from herbivory to carnivory or carnivory to herbivory would be rare except via an intermediate stage of omnivory.


Omnivorous mammals

Various
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s are omnivorous in the wild, such as species of hominids, pigs, badgers, bears, foxes,
coati Coatis, also known as coatimundis (), are members of the family Procyonidae in the genera ''Nasua'' and ''Nasuella''. They are diurnal mammals native to South America, Central America, Mexico, and the southwestern United States. The name ...
s, civets, hedgehogs, opossums, skunks,
sloth Sloths are a group of Neotropical xenarthran mammals constituting the suborder Folivora, including the extant arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their l ...
s, squirrels, raccoons, chipmunks, mice, hamsters and rats. Most bear species are omnivores, but individual diets can range from almost exclusively herbivorous ( hypocarnivore) to almost exclusively carnivorous ( hypercarnivore), depending on what food sources are available locally and seasonally. Polar bears are classified as carnivores, both taxonomically (they are in the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...
Carnivora), and behaviorally (they subsist on a largely carnivorous diet). Depending on the species of bear, there is generally a preference for one class of food, as plants and animals are digested differently. Canines including wolves, dogs, dingoes, and
coyote The coyote (''Canis latrans'') is a species of canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological nich ...
s eat some plant matter, but they have a general preference and are evolutionarily geared towards meat. However, the maned wolf is a canid whose diet is naturally 50% plant matter. Like most arboreal species, squirrels are primarily granivores, subsisting on nuts and seeds. However, like virtually all mammals, squirrels avidly consume some animal food when it becomes available. For example, the American eastern gray squirrel has been introduced to parts of Britain, continental Europe and South Africa. Its effect on populations of nesting birds is often serious because of consumption of eggs and nestlings.


Other species

Various
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s are omnivorous, with diets varying from
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, ras ...
and nectar to
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s,
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wor ...
s,
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
, and small
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s. Examples include cranes, cassowaries,
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adu ...
s, crows and related corvids, kea,
rallidae The rails, or Rallidae, are a large cosmopolitan family of small- to medium-sized, ground-living birds. The family exhibits considerable diversity and includes the crakes, coots, and gallinules. Many species are associated with wetlands, alth ...
, and
rheas The rheas ( ), also known as ñandus ( ) or South American ostriches, are large ratites (flightless birds without a keel on their sternum bone) in the order Rheiformes, native to South America, distantly related to the ostrich and emu. Most tax ...
. In addition, some lizards (such as Galapagos Lava Lizard),
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked t ...
s, fish (such as
piranha A piranha or piraña (, , or ; or , ) is one of a number of freshwater fish in the family Serrasalmidae, or the subfamily Serrasalminae within the tetra family, Characidae in order Characiformes. These fish inhabit South American rivers, f ...
s and
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
), and
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
s are omnivorous. Quite often, mainly herbivorous creatures will eagerly eat small quantities of animal food when it becomes available. Although this is trivial most of the time, omnivorous or herbivorous birds, such as sparrows, often will feed their chicks insects while food is most needed for growth. On close inspection it appears that nectar-feeding birds such as sunbirds rely on the ants and other insects that they find in flowers, not for a richer supply of protein, but for essential nutrients such as
cobalt Cobalt is a chemical element with the symbol Co and atomic number 27. As with nickel, cobalt is found in the Earth's crust only in a chemically combined form, save for small deposits found in alloys of natural meteoric iron. The free element, p ...
/ vitamin b12 that are absent from nectar. Similarly, monkeys of many species eat maggoty fruit, sometimes in clear preference to sound fruit.Ewing, Jack (2005). ''Monkeys Are Made of Chocolate''. Publisher: Pixyjack Press. . When to refer to such animals as omnivorous, or otherwise, is a question of context and emphasis, rather than of definition.


See also


References

{{Authority control Animals by eating behaviors Ethology