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A nose is a protuberance in vertebrates that houses the
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called tur ...
s, or nares, which receive and expel air for respiration alongside the mouth. Behind the nose are the
olfactory mucosa The olfactory mucosa is located in the upper region of the nasal cavity and is made up of the olfactory epithelium and the underlying lamina propria, connective tissue containing fibroblasts, blood vessels, Bowman's glands and bundles of fine axo ...
and the sinuses. Behind the nasal cavity, air next passes through the pharynx, shared with the digestive system, and then into the rest of the respiratory system. In humans, the nose is located centrally on the face and serves as an alternative respiratory passage especially during suckling for infants. The protruding nose that completely separate from the mouth part is a characteristic found only in
therian mammals Theria (; Greek: , wild beast) is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the placental mammals) and the metatherians (including the marsupials) but excludes the egg-laying monotremes. Chara ...
. It has been theorized that this unique mammalian nose evolved from the anterior part of the upper jaw of the reptilian-like ancestors ( synapsids).


Air treatment

Acting as the first interface between the external environment and an animal's delicate internal lungs, a nose conditions incoming air, both as a function of thermal regulation and filtration during respiration, as well as enabling the sensory perception of smell. Hair inside nostrils filter incoming air, as a first line of defense against dust particles, smoke, and other potential obstructions that would otherwise inhibit respiration, and as a kind of filter against airborne illness. In addition to acting as a filter, mucus produced within the nose supplements the body's effort to maintain temperature, as well as contributes moisture to integral components of the respiratory system. Capillary structures of the nose warm and humidify air entering the body; later, this role in retaining moisture enables conditions for alveoli to properly exchange O2 for CO2 (i.e., respiration) within the lungs. During exhalation, the capillaries then aid recovery of some moisture, mostly as a function of thermal regulation, again.


Sense of direction

The wet nose of dogs is useful for the perception of direction. The sensitive cold receptors in the skin detect the place where the nose is cooled the most and this is the direction a particular smell that the animal just picked up comes from.


Structure in air-breathing forms

In
amphibian Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arborea ...
s and lungfish, the nostrils open into small sacs that, in turn, open into the forward roof of the mouth through the choanae. These sacs contain a small amount of olfactory epithelium, which, in the case of
caecilian Caecilians (; ) are a group of limbless, vermiform or serpentine amphibians. They mostly live hidden in the ground and in stream substrates, making them the least familiar order of amphibians. Caecilians are mostly distributed in the tropics of ...
s, also lines a number of neighbouring tentacles. Despite the general similarity in structure to those of amphibians, the nostrils of lungfish are not used in respiration, since these animals breathe through their mouths. Amphibians also have a vomeronasal organ, lined by olfactory epithelium, but, unlike those of
amniote Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are distin ...
s, this is generally a simple sac that, except in salamanders, has little connection with the rest of the nasal system. In reptiles, the nasal chamber is generally larger, with the choanae located much further back in the roof of the mouth. In
crocodilian Crocodilia (or Crocodylia, both ) is an Order (biology), order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous Period (geology), period (Cenomanian St ...
s, the chamber is exceptionally long, helping the animal to breathe while partially submerged. The reptilian nasal chamber is divided into three parts: an anterior ''vestibule'', the main olfactory chamber, and a posterior nasopharynx. The olfactory chamber is lined by olfactory epithelium on its upper surface and possesses a number of turbinates to increase the sensory area. The vomeronasal organ is well-developed in lizards and snakes, in which it no longer connects with the nasal cavity, opening directly into the roof of the mouth. It is smaller in turtles, in which it retains its original nasal connection, and is absent in adult crocodilians. Birds have a similar nose to reptiles, with the nostrils located at the upper rear part of the beak. Since they generally have a poor sense of smell, the olfactory chamber is small, although it does contain three turbinates, which sometimes have a complex structure similar to that of mammals. In many birds, including doves and
fowl Fowl are birds belonging to one of two biological orders, namely the gamefowl or landfowl (Galliformes) and the waterfowl (Anseriformes). Anatomical and molecular similarities suggest these two groups are close evolutionary relatives; together ...
s, the nostrils are covered by a horny protective shield. The vomeronasal organ of birds is either under-developed or altogether absent, depending on the species. The nasal cavities in mammals are both fused into one. Among most species they are exceptionally large, typically occupying up to half the length of the skull. In some groups, however, including primates,
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most bi ...
s, and
cetaceans Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...
, the nose has been secondarily reduced, and these animals consequently have a relatively poor sense of smell. The nasal cavity of mammals has been enlarged, in part, by the development of a
palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ...
cutting off the entire upper surface of the original oral cavity, which consequently becomes part of the nose, leaving the palate as the new roof of the mouth. The enlarged nasal cavity contains complex turbinates forming coiled scroll-like shapes that help to warm the air before it reaches the lungs. The cavity also extends into neighbouring skull bones, forming additional air cavities known as paranasal sinuses. In
cetacean Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel the ...
s, the nose has been reduced to one or two blowholes, which are the nostrils that have migrated to the top of the head. This adaptation gave cetaceans a more streamlined body shape and the ability to breathe while mostly submerged. Conversely, the
elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae a ...
's nose has elaborated into a long, muscular, manipulative organ called the
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...
. The vomeronasal organ of mammals is generally similar to that of reptiles. In most species, it is located in the floor of the nasal cavity, and opens into the mouth via two ''nasopalatine ducts'' running through the palate, but it opens directly into the nose in many rodents. It is, however, lost in bats, and in many primates, including humans.


In fish

Fish have a relatively good sense of smell. Unlike that of tetrapods, the nose has no connection with the mouth, nor any role in respiration. Instead, it generally consists of a pair of small pouches located behind the nostrils at the front or sides of the head. In many cases, each of the nostrils is divided into two by a fold of skin, allowing water to flow into the nose through one side and out through the other. The pouches are lined by olfactory epithelium, and commonly include a series of internal folds to increase the surface area, often forming an elaborate "olfactory rosette". In some teleosts, the pouches branch off into additional sinus-like cavities, while in
coelacanth The coelacanths ( ) are fish belonging to the order Actinistia that includes two extant species in the genus ''Latimeria'': the West Indian Ocean coelacanth (''Latimeria chalumnae''), primarily found near the Comoro Islands off the east coast ...
s, they form a series of tubes. In the earliest vertebrates, there was only one nostril and olfactory pouch, and the nasal passage was connected to the hypophysis. The same anatomy is observed in the most primitive living vertebrates, the
lamprey Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like ...
s and hagfish. In gnathostome ancestors, the olfactory apparatus gradually became paired (presumably to allow sense of direction of smells), and freeing the midline from the nasal passage allowed evolution of jaws.


See also

*
Human nose The human nose is the most protruding part of the face. It bears the nostrils and is the first organ of the respiratory system. It is also the principal organ in the olfactory system. The shape of the nose is determined by the nasal bones ...
* Nasal bridge *
Obligate nasal breathing Obligate nasal breathing describes a physiological necessity to breathe through the nose (or other forms of external nares, depending on the species) as opposed to breathing through the mouth. Definition The term may be misleading, as it implie ...
*
Rhinarium The rhinarium (New Latin, "belonging to the nose"; plural: rhinaria) is the furless skin surface surrounding the external openings of the nostrils in many mammals. Commonly it is referred to as the tip of the '' snout'', and breeders of cats and ...
, the wet, naked surface around the nostrils in most mammals, absent in
haplorrhine Haplorhini (), the haplorhines (Greek for "simple-nosed") or the "dry-nosed" primates, is a suborder of primates containing the tarsiers and the simians (Simiiformes or anthropoids), as sister of the Strepsirrhini ("moist-nosed"). The name is some ...
primates such as humans


References


External links

* * {{Authority control Human head and neck Respiratory system Olfactory system Facial features