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Morphology is a branch of
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...
dealing with the study of the form and structure of
organism In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and ...
s and their specific structural features. This includes aspects of the outward appearance ( shape, structure, colour,
pattern A pattern is a regularity in the world, in human-made design, or in abstract ideas. As such, the elements of a pattern repeat in a predictable manner. A geometric pattern is a kind of pattern formed of geometric shapes and typically repeated li ...
, size), i.e. external morphology (or eidonomy), as well as the form and structure of the internal parts like
bones A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, a ...
and organs, i.e. internal morphology (or anatomy). This is in contrast to physiology, which deals primarily with function. Morphology is a branch of life science dealing with the study of gross structure of an
organism In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and ...
or taxon and its component parts.


History

The
etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...
of the word "morphology" is from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
(), meaning "form", and (), meaning "word, study, research". While the concept of form in biology, opposed to
function Function or functionality may refer to: Computing * Function key, a type of key on computer keyboards * Function model, a structured representation of processes in a system * Function object or functor or functionoid, a concept of object-oriente ...
, dates back to Aristotle (see
Aristotle's biology Aristotle's biology is the theory of biology, grounded in systematic observation and collection of data, mainly zoological, embodied in Aristotle's books on the science. Many of his observations were made during his stay on the island of Le ...
), the field of morphology was developed by Johann Wolfgang von Goethe (1790) and independently by the German anatomist and physiologist Karl Friedrich Burdach (1800). Among other important theorists of morphology are Lorenz Oken, Georges Cuvier, Étienne Geoffroy Saint-Hilaire, Richard Owen,
Karl Gegenbaur Karl Gegenbaur (21 August 1826 – 14 June 1903)"Karl Gegenbaur – Encyclopædia Britannica" (biography), ''Encyclopædia Britannica'', 2006, Britannica.coBritannica-KarlG was a German anatomist and professor who demonstrated that the field of c ...
and Ernst Haeckel. In 1830, Cuvier and E.G.Saint-Hilaire engaged in a famous debate, which is said to exemplify the two major deviations in biological thinking at the time – whether animal structure was due to function or evolution.


Divisions of morphology

*
Comparative morphology Comparative anatomy is the study of similarities and differences in the anatomy of different species. It is closely related to evolutionary biology and phylogeny (the evolution of species). The science began in the classical era, continuing i ...
is analysis of the patterns of the locus of structures within the body plan of an organism, and forms the basis of taxonomical categorization. *
Functional morphology Functional may refer to: * Movements in architecture: ** Functionalism (architecture) ** Form follows function * Functional group, combination of atoms within molecules * Medical conditions without currently visible organic basis: ** Functional sy ...
is the study of the relationship between the structure and function of morphological features. * Experimental morphology is the study of the effects of external factors upon the morphology of organisms under experimental conditions, such as the effect of genetic mutation. * Anatomy is a "branch of morphology that deals with the structure of organisms". * Molecular morphology is a rarely used term, usually referring to the superstructure of polymers such as fiber formation or to larger composite assemblies. The term is commonly not applied to the spatial structure of individual molecules. * Gross morphology refers to the collective structures of an organism as a whole as a general description of the form and structure of an organism, taking into account all of its structures without specifying an individual structure.


Morphology and classification

Most taxa differ morphologically from other taxa. Typically, closely related taxa differ much less than more distantly related ones, but there are exceptions to this. Cryptic species are
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
which look very similar, or perhaps even outwardly identical, but are reproductively isolated. Conversely, sometimes unrelated taxa acquire a similar appearance as a result of convergent evolution or even mimicry. In addition, there can be morphological differences within a species, such as in ''
Apoica flavissima ''Apoica flavissima'' is a paper wasp found primarily in South America. The species is distinguishable by its light coloring, unique single comb nests, and nocturnal nature. A notable feature of this species is the size dimorphism between queens ...
'' where queens are significantly smaller than workers. A further problem with relying on morphological data is that what may appear, morphologically speaking, to be two distinct species, may in fact be shown by DNA analysis to be a single species. The significance of these differences can be examined through the use of allometric engineering in which one or both species are manipulated to phenocopy the other species. A step relevant to the evaluation of morphology between traits/features within species, includes an assessment of the terms: homology an
homoplasy
Homology between features indicate that those features have been derived from a common ancestor. Alternatively, homoplasy between features describes those that can resemble each other, but derive independently via parallel or convergent evolution.


3D cell morphology: classification

Invention and development of microscopy enable the observation of 3-D cell morphology with both high spatial and temporal resolution. The dynamic processes of these cell morphology which are controlled by a complex system play an important role in varied important biological process, such as immune and invasive responses.


See also

* Comparative anatomy *
Computational anatomy Computational anatomy is an interdisciplinary field of biology focused on quantitative investigation and modelling of anatomical shapes variability. It involves the development and application of mathematical, statistical and data-analytical metho ...
* Insect morphology *
Morphometrics Morphometrics (from Greek μορϕή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
* Neuromorphology *
Phenetics In biology, phenetics ( el, phainein – to appear) , also known as taximetrics, is an attempt to classify organisms based on overall similarity, usually in morphology or other observable traits, regardless of their phylogeny or evolutionary re ...
* Phenotype *
Phenotypic plasticity Phenotypic plasticity refers to some of the changes in an organism's behavior, morphology and physiology in response to a unique environment. Fundamental to the way in which organisms cope with environmental variation, phenotypic plasticity encompa ...
*
Plant morphology Phytomorphology is the study of the physical form and external structure of plants.Raven, P. H., R. F. Evert, & S. E. Eichhorn. ''Biology of Plants'', 7th ed., page 9. (New York: W. H. Freeman, 2005). . This is usually considered distinct from ...


References


External links

* {{Authority control Branches of biology Comparative anatomy