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The metatarsal bones, or metatarsus, are a group of five
long bone The long bones are those that are longer than they are wide. They are one of five types of bones: long, short, flat, irregular and sesamoid. Long bones, especially the femur and tibia, are subjected to most of the load during daily activities ...
s in the
foot The foot ( : feet) is an anatomical structure found in many vertebrates. It is the terminal portion of a limb which bears weight and allows locomotion. In many animals with feet, the foot is a separate organ at the terminal part of the leg mad ...
, located between the tarsal bones of the hind- and mid-foot and the
phalanges The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. ...
of the toes. Lacking individual names, the metatarsal bones are numbered from the medial side (the side of the great toe): the first,
second The second (symbol: s) is the unit of time in the International System of Units (SI), historically defined as of a day – this factor derived from the division of the day first into 24 hours, then to 60 minutes and finally to 60 seconds ea ...
,
third Third or 3rd may refer to: Numbers * 3rd, the ordinal form of the cardinal number 3 * , a fraction of one third * 1⁄60 of a ''second'', or 1⁄3600 of a ''minute'' Places * 3rd Street (disambiguation) * Third Avenue (disambiguation) * Hi ...
, fourth, and
fifth metatarsal The fifth metatarsal bone is a long bone in the foot, and is palpable along the distal outer edges of the feet. It is the second smallest of the five metatarsal bones. The fifth metatarsal is analogous to the fifth metacarpal bone in the hand. As ...
(often depicted with
Roman numerals Roman numerals are a numeral system that originated in ancient Rome and remained the usual way of writing numbers throughout Europe well into the Late Middle Ages. Numbers are written with combinations of letters from the Latin alphabet, ...
). The metatarsals are analogous to the
metacarpal In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ar ...
bones of the
hand A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the koala (which has two opposable thumbs on each "h ...
. The lengths of the metatarsal bones in humans are, in descending order, second, third, fourth, fifth, and first.


Structure

The five metatarsals are
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...
convex long bones consisting of a shaft or body, a base (
proximal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position ...
ly), and a head (
distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
ly).Platzer 2004, p. 220 The body is prismoid in form, tapers gradually from the tarsal to the phalangeal extremity, and is curved longitudinally, so as to be concave below, slightly convex above. The base or posterior extremity is wedge-shaped, articulating proximally with the tarsal bones, and by its sides with the contiguous metatarsal bones: its dorsal and plantar surfaces are rough for the attachment of
ligament A ligament is the fibrous connective tissue that connects bones to other bones. It is also known as ''articular ligament'', ''articular larua'', ''fibrous ligament'', or ''true ligament''. Other ligaments in the body include the: * Peritoneal l ...
s. The head or distal extremity presents a convex articular surface, oblong from above downward, and extending farther backward below than above. Its sides are flattened, and on each is a depression, surmounted by a
tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection ...
, for ligamentous attachment. Its plantar surface is grooved antero-posteriorly for the passage of the flexor
tendon A tendon or sinew is a tough, high-tensile-strength band of dense fibrous connective tissue that connects muscle to bone. It is able to transmit the mechanical forces of muscle contraction to the skeletal system without sacrificing its ability ...
s, and marked on either side by an articular eminence continuous with the terminal articular surface.Gray's 1918, ''6d. 2. The Metatarsus'' During growth, the growth plates are located distally on the metatarsals, except on the first metatarsal where it is located proximally. Yet it is quite common to have an accessory growth plate on the distal first metatarsal.


Articulations

The base of each metatarsal bone articulates with one or more of the tarsal bones at the
tarsometatarsal joint The tarsometatarsal joints (Lisfranc joints) are arthrodial joints in the foot. The tarsometatarsal joints involve the first, second and third cuneiform bones, the cuboid bone and the metatarsal bones. The eponym of Lisfranc joint is 18th-19th ce ...
s, and the head with one of the first row of phalanges at the
metatarsophalangeal joint The metatarsophalangeal joints (MTP joints), also informally known as toe knuckles, are the joints between the metatarsal bones of the foot and the proximal bones ( proximal phalanges) of the toes. They are condyloid joints, meaning that an elli ...
s. Their bases also articulate with each other at the intermetatarsal joints * The ''first'' metatarsal articulates with the medial cuneiform, and to a small extent to the intermediate cuneiform.Platzer 2004, p. 218 * the ''second'' with all three cuneiforms. * the ''third'' with the lateral cuneiform. * the ''fourth'' with the lateral cuneiform and the
cuboid In geometry, a cuboid is a hexahedron, a six-faced solid. Its faces are quadrilaterals. Cuboid means "like a cube", in the sense that by adjusting the length of the edges or the angles between edges and faces a cuboid can be transformed into a c ...
. * The ''fifth'' with the cuboid.


Muscle attachments


Clinical significance


Injuries

The metatarsal bones are often broken by
association football Association football, more commonly known as football or soccer, is a team sport played between two teams of 11 players who primarily use their feet to propel the ball around a rectangular field called a pitch. The objective of the game is ...
players. These and other recent cases have been attributed to the lightweight design of modern football boots, which provide less protection to the foot. In 2010 some soccer players began testing a new sock that incorporated a rubber silicone pad over the foot to provide protection to the top of the foot. Stress fractures are thought to account for 16% of injuries related to sports participation, and the metatarsals are the bones most often involved. These fractures are sometimes called
march fracture March fracture, is the fracture of the distal third of one of the metatarsals occurring because of recurrent stress. It is more common in soldiers, but also occurs in hikers, organists, and people whose duties entail much standing (such as hospit ...
s, based on their traditional association with military recruits after long marches. The second and third metatarsals are fixed while walking, thus these metatarsals are common sites of injury. The fifth metatarsal may be fractured if the foot is oversupinated during locomotion. Protection from injuries can be given by the use of safety footwear which can use built-in or removable metatarsal guards.


Additional images

Image:Footx.jpg, X-ray of foot. Image:Gray290 - Mratatarsus.png, Skeleton of left foot. Medial aspect. Image:Gray360.png, Oblique section of left intertarsal and tarsometatarsal articulations, showing the synovial cavities. File:Slide2xzxzx.JPG, Ankle and tarsometarsal joints, showing bones of foot. Deep dissection. File:Nitti MetatarsalGuard.jpg, alt=Nitti Safety Footwear with removable metatarsal guard., Safety footwear with removable metatarsal guard.


See also

*
Arches of the foot The arches of the foot, formed by the tarsal and metatarsal bones, strengthened by ligaments and tendons, allow the foot to support the weight of the body in the erect posture with the least weight. They are categorized as longitudinal and tran ...
*
Ball (foot) The ball of the foot is the padded portion of the sole between the toes and the arch, underneath the heads of the metatarsal bones. In comparative foot morphology, the ball is most analogous to the metacarpal (forepaw) or metatarsal (hindpaw) ...
* Bone terminology *
Terms for anatomical location Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
*
Jones fracture A Jones fracture is a broken bone in a specific part of the fifth metatarsal of the foot between the base and middle part that is known for its high rate of delayed healing or nonunion. It results in pain near the midportion of the foot on the ...
*
Lisfranc injury A Lisfranc injury, also known as Lisfranc fracture, is an injury of the foot in which one or more of the metatarsal bones are displaced from the tarsus. The injury is named after Jacques Lisfranc de St. Martin, a French surgeon and gynecolog ...


Notes


References

* *


External links

* {{Authority control Bones of the lower limb Bones of the foot