malleus
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The malleus, or hammer, is a hammer-shaped small bone or
ossicle The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditory ...
of the
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in the ...
. It connects with the
incus The ''incus'' (plural incudes) or anvil is a bone in the middle ear. The anvil-shaped small bone is one of three ossicles in the middle ear. The ''incus'' receives vibrations from the ''malleus'', to which it is connected laterally, and transmit ...
, and is attached to the inner surface of the
eardrum In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped membrane that separates the external ear The outer ear, external ear, or auris externa is the extern ...
. The word is Latin for 'hammer' or 'mallet'. It transmits the sound vibrations from the
eardrum In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped membrane that separates the external ear The outer ear, external ear, or auris externa is the extern ...
to the ''
incus The ''incus'' (plural incudes) or anvil is a bone in the middle ear. The anvil-shaped small bone is one of three ossicles in the middle ear. The ''incus'' receives vibrations from the ''malleus'', to which it is connected laterally, and transmit ...
'' (anvil).


Structure

The malleus is a bone situated in the middle ear. It is the first of the three ossicles, and attached to the
tympanic membrane In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped membrane that separates the external ear from the middle ear. Its function is to transmit sound from the a ...
. The head of the malleus is the large protruding section, which attaches to the
incus The ''incus'' (plural incudes) or anvil is a bone in the middle ear. The anvil-shaped small bone is one of three ossicles in the middle ear. The ''incus'' receives vibrations from the ''malleus'', to which it is connected laterally, and transmit ...
. The head connects to the neck of malleus. The bone continues as the handle (or manubrium) of malleus, which connects to the tympanic membrane. Between the neck and handle of the malleus, lateral and anterior processes emerge from the bone. The bone is oriented so that the head is superior and the handle is inferior.


Development

Embryologically, the malleus is derived from the first
pharyngeal arch The pharyngeal arches, also known as visceral arches'','' are structures seen in the embryonic development of vertebrates that are recognisable precursors for many structures. In fish, the arches are known as the branchial arches, or gill arche ...
along with the ''
incus The ''incus'' (plural incudes) or anvil is a bone in the middle ear. The anvil-shaped small bone is one of three ossicles in the middle ear. The ''incus'' receives vibrations from the ''malleus'', to which it is connected laterally, and transmit ...
''. It grows from
Meckel's cartilage In humans, the cartilaginous bar of the mandibular arch is formed by what are known as Meckel's cartilages (right and left) also known as Meckelian cartilages; above this the incus and malleus are developed. Meckel's cartilage arises from the fir ...
.


Function

The malleus is one of three ossicles in the
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in the ...
which transmit sound from the tympanic membrane (ear drum) to the inner ear. The malleus receives vibrations from the tympanic membrane and transmits this to the incus.


Clinical significance

The malleus may be
palpated Palpation is the process of using one's hands to check the body, especially while perceiving/diagnosing a disease or illness. Usually performed by a health care practitioner, it is the process of feeling an object in or on the body to determine ...
by
surgeons In modern medicine, a surgeon is a medical professional who performs surgery. Although there are different traditions in different times and places, a modern surgeon usually is also a licensed physician or received the same medical training as ...
during
ear surgery Otorhinolaryngology ( , abbreviated ORL and also known as otolaryngology, otolaryngology–head and neck surgery (ORL–H&N or OHNS), or ear, nose, and throat (ENT)) is a surgical subspeciality within medicine that deals with the surgical a ...
. It may become fixed in place due to surgical complications, causing hearing loss. This may be corrected with further surgery.


History

Several sources attribute the discovery of the malleus to the
anatomist Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
and
philosopher A philosopher is a person who practices or investigates philosophy. The term ''philosopher'' comes from the grc, φιλόσοφος, , translit=philosophos, meaning 'lover of wisdom'. The coining of the term has been attributed to the Greek th ...
Alessandro Achillini Alessandro Achillini (''Latin'' Alexander Achillinus; 20 or 29 October 1463 (or possibly 1461)2 August 1512) was an Italian philosopher and physician. He is known for the anatomic studies that he was able to publish, made possible by a 13th-cent ...
. The first brief written description of the malleus was by
Berengario da Carpi Jacopo Berengario da Carpi (also known as Jacobus Berengarius Carpensis, Jacopo Barigazzi, Giacomo Berengario da Carpi or simply Carpus; c. 1460 – c. 1530) was an Italian physician. His book "''Isagoge breves''" published in 1522 made him the mo ...
in his ''Commentaria super anatomia Mundini'' (1521). Niccolo Massa's ''Liber introductorius anatomiae'' described the malleus in slightly more detail and likened both it and the
incus The ''incus'' (plural incudes) or anvil is a bone in the middle ear. The anvil-shaped small bone is one of three ossicles in the middle ear. The ''incus'' receives vibrations from the ''malleus'', to which it is connected laterally, and transmit ...
to little hammers terming them ''malleoli''.


Other animals

The ''malleus'' is unique to mammals, and evolved from a lower jaw bone in basal amniotes called the
articular The articular bone is part of the lower jaw of most vertebrates, including most jawed fish, amphibians, birds and various kinds of reptiles, as well as ancestral mammals. Anatomy In most vertebrates, the articular bone is connected to two oth ...
, which still forms part of the jaw joint in reptiles and birds.


Additional images


See also

* Bone terminology *
Evolution of mammalian auditory ossicles The evolution of mammalian auditory ossicles was an evolutionary event that resulted in the formation of the bones of the mammalian middle ear. These bones, or ossicles, are a defining characteristic of all mammals. The event is well-documented ...
*
Ligaments of malleus The ligaments of malleus are three ligaments that attach the malleus in the middle ear. They are the anterior, lateral and superior ligaments. The anterior ligament of the malleus also known as Casserio's ligament is a fibrous band that extends ...
*
Terms for anatomical location Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...


References

{{Authority control Auditory system Bones of the head and neck Ear Ossicles