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The longleaf pine (''Pinus palustris'') is a
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...
species native to the
Southeastern United States The Southeastern United States, also referred to as the American Southeast or simply the Southeast, is a geographical List of regions in the United States, region of the United States. It is located broadly on the eastern portion of the south ...
, found along the coastal plain from
East Texas East Texas is a broadly defined cultural, geographic, and ecological region in the eastern part of the U.S. state of Texas that comprises most of 41 counties. It is primarily divided into Northeast and Southeast Texas. Most of the region co ...
to southern
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth are ...
, extending into northern and central
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
. In this area it is also known as "yellow pine" or "long leaf yellow pine", although it is properly just one out of a number of species termed yellow pine. It reaches a height of and a diameter of . In the past, before extensive logging, they reportedly grew to with a diameter of . The tree is a cultural symbol of the
Southern United States The Southern United States (sometimes Dixie, also referred to as the Southern States, the American South, the Southland, or simply the South) is a geographic and cultural region of the United States of America. It is between the Atlantic Ocean ...
, being the official state tree of
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
. Contrary to popular belief, this particular species of pine is not officially the state tree of
North Carolina North Carolina () is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. The state is the List of U.S. states and territories by area, 28th largest and List of states and territories of the United ...
.


Description

The bark is thick, reddish-brown, and scaly. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are dark green and needle-like, and occur in bundles of mainly three, sometimes two or four, especially in seedlings. They often are twisted and in length. A local race of ''P. palustris'' in a cove near Rockingham, North Carolina, have needles up to 24 inches (61 centimeters ) in length. It is one of the two Southeastern U.S. pines with long needles, the other being slash pine. The cones, both female seed cones (ovulate strobili) and male pollen cones (staminate strobili), are initiated during the growing season before buds emerge. Pollen cones begin forming in their buds in July, while seed conelets are formed during a relatively short period of time in August. Pollination occurs early the following spring, with the male cones long. The female (seed) cones mature in about 20 months from pollination; when mature, they are yellow-brown in color, long, and broad, opening to , and have a small, but sharp, downward-pointing spine on the middle of each scale. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s are long, with a wing. Longleaf pine takes 100 to 150 years to become full size and may live to be 500 years old. When young, they grow a long taproot, which usually is long; by maturity, they have a wide spreading lateral root system with several deep 'sinker' roots. They grow on well-drained, usually sandy
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
, characteristically in pure stands. Longleaf pine also is known as being one of several species grouped as a southern yellow pine or longleaf yellow pine, and in the past as pitch pine (a name dropped as it caused confusion with pitch pine, '' Pinus rigida'').


Etymology

The species
epithet An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...
''palustris'' is Latin for "of the marsh" and indicates its common habitat. The scientific name meaning "of marshes" is a misunderstanding on the part of Philip Miller, who described the species, after seeing longleaf pine forests with temporary winter flooding.


Ecology

Longleaf pine is highly pyrophytic (resistant to
wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identi ...
) and dependent on fire. Their thick bark and growth habits help to provide a tolerance to fire. Periodic natural wildfire and anthropogenic fires select for this species by removing competition and exposing bare soil for successful germination of seeds. The lack of medium-tall trees (called a midstory canopy) leads to open longleaf pine
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s or savannas. New seedlings do not appear at all tree-like and resemble a dark-green fountain of needles. This form is called the grass stage. During this stage, which lasts for 5–12 years, vertical growth is very slow, and the tree may take a number of years simply to grow ankle high. After that, it has a growth spurt, especially if it is in a gap or no tree canopy is above it. In the grass stage, it is very resistant to low intensity fires because the terminal bud is protected from lethal heating by the tightly packed needles. While relatively immune to fire at this stage, the plant is quite appealing to feral pigs; the early settlers' habit of releasing swine into the woodlands to feed may have been partly responsible for the decline of the species. Longleaf pine forests are rich in
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity'') ...
. They are well-documented for their high levels of plant diversity, in groups including sedges, grasses, carnivorous plants, and orchids. These forests also provide habitat for
gopher tortoise The gopher tortoise (''Gopherus polyphemus'') is a species of tortoise in the family Testudinidae. The species is native to the southeastern United States. The gopher tortoise is seen as a keystone species because it digs burrows that provid ...
s, which as keystone species, dig burrows that provide habitat for hundreds of other species of animals. The red-cockaded woodpecker is dependent on mature pine forests and is now
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
as a result of this decline. Longleaf pine
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s are large and nutritious, forming a significant food source for
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s (notably the
brown-headed nuthatch The brown-headed nuthatch (''Sitta pusilla'') is a small songbird endemic to pine forests throughout the Southeastern United States. Genetic analyses indicated low differentiation between northern and southern populations in Florida, but the stud ...
) and other
wildlife Wildlife refers to undomesticated animal species, but has come to include all organisms that grow or live wild in an area without being introduced by humans. Wildlife was also synonymous to game: those birds and mammals that were hunted ...
. Nine
salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All t ...
species and 26
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
species are characteristic of pine savannas, along with 56 species of
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalia ...
s, 13 of which could be considered specialists on this habitat. The Red Hills Region of Florida and Georgia is home to some of the best-preserved stands of longleaf pines. These forests have been burned regularly for many decades to encourage bobwhite quail habitat in private hunting plantations.


Native range, restoration, and protection

Before European settlement, longleaf pine forest dominated as much as stretching from Virginia south to Florida and west to East Texas. Its range was defined by the frequent widespread fires that were lit by humans and occurred naturally throughout the southeast. In the late 19th century, these virgin timber stands were "among the most sought-after timber trees in the country." This rich ecosystem now has been relegated to less than 5% of its presettlement range due to fire suppression and clear-cutting practices:
As they stripped the woods of their trees, loggers left mounds of flammable debris that frequently fueled catastrophic fires, destroying both the remaining trees and seedlings. The exposed earth left behind by clear-cutting operations was highly susceptible to erosion, and nutrients were washed from the already porous soils. This further destroyed the natural seeding process. At the peak of the timber cutting in the 1890s and first decade of the new century, the longleaf pine forests of the Sandhills were providing millions of board feet of timber each year. The timber cutters gradually moved across the South; by the 1920s, most of the "limitless" virgin longleaf pine forests were gone. :— Jerry Simmons, "ASLC Large Operation from Beginnings"
In " pine barrens" most of the day. Low, level, sandy tracts; the pines wide apart; the sunny spaces between full of beautiful abounding grasses,
liatris ''Liatris'' (), commonly known as gayfeather and blazing star. is a genus of flowering plants in the tribe Eupatorieae within the family Asteraceae native to North America (Canada, United States, Mexico and the Bahamas). Some species are used ...
, long, wand-like solidago, saw palmettos, etc., covering the ground in garden style. Here I sauntered in delightful freedom, meeting none of the cat-clawed vines, or shrubs, of the alluvial bottoms. : –
John Muir John Muir ( ; April 21, 1838December 24, 1914), also known as "John of the Mountains" and "Father of the National Parks", was an influential Scottish-American naturalist, author, environmental philosopher, botanist, zoologist, glaciologis ...
Efforts are being made to restore longleaf pine ecosystems within its natural range. Some groups such as the Longleaf Alliance are actively promoting research, education, and management of the longleaf pine. The
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of comme ...
offers cost-sharing and technical assistance to private landowners for longleaf restoration through the NRCS Longleaf Pine Initiative. Similar programs are available through most state forestry agencies in the longleaf's native range. In August 2009, the Alabama Forestry Commission received $1.757 million in stimulus money to restore longleaf pines in state forests. Four large core areas within the range of the species provide the opportunity to protect the biological diversity of the coastal plain and to restore wilderness areas east of the Mississippi River. Each of these four (
Eglin Air Force Base Eglin Air Force Base is a United States Air Force (USAF) base in the western Florida Panhandle, located about southwest of Valparaiso in Okaloosa County. The host unit at Eglin is the 96th Test Wing (formerly the 96th Air Base Wing). The 9 ...
: 187,000+ ha; Apalachicola National Forest: 228,000+ ha;
Okefenokee The Okefenokee Swamp is a shallow, 438,000-acre (177,000 ha), peat-filled wetland straddling the Georgia– Florida line in the United States. A majority of the swamp is protected by the Okefenokee National Wildlife Refuge and the Okefeno ...
- Osceola: 289,000+ ha; De Soto National Forest: 200,000+ ha) have nearby lands that offer the potential to expand the total protected territory for each area to well beyond 500,000 ha. These areas would provide the opportunity not only to restore forest stands, but also to restore populations of native plants and animals threatened by landscape fragmentation. Notable eccentric populations exist within the Uwharrie National Forest in the central Piedmont region of North Carolina. These have survived owing to relative inaccessibility, and in one instance, intentional protection in the 20th century by a private landowner (a property now owned and conserved by the LandTrust for Central North Carolina). The United States Forest Service is conducting prescribed burning programs in the 258,864-acre Francis Marion National Forest, located outside of
Charleston, South Carolina Charleston is the largest city in the U.S. state of South Carolina, the county seat of Charleston County, and the principal city in the Charleston–North Charleston metropolitan area. The city lies just south of the geographical midpoint o ...
. They are hoping to increase the longleaf pine forest type to by 2017 and in the long term. In addition to longleaf restoration, prescribed burning will enhance the endangered red-cockaded woodpeckers' preferred habitat of open, park-like stands, provide habitat for wildlife dependent on grass-shrub habitat, which is very limited, and reduce the risk of damaging wildfires. Since the 1960s, longleaf restoration has been ongoing on almost 95,000 acres of state and federal land in the sandhills region of South Carolina, between the
piedmont it, Piemontese , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = , demographics1_title2 ...
and coastal plain. The region is characterized by deep, infertile sands deposited by a prehistoric sea, with generally arid conditions. By the 1930s, most of the native longleaf had been logged, and the land was heavily eroded. Between 1935 and 1939, the federal government purchased large portions of this area from local landowners as a relief measure under the
Resettlement Administration The Resettlement Administration (RA) was a New Deal U.S. federal agency created May 1, 1935. It relocated struggling urban and rural families to communities planned by the federal government. On September 1, 1937, it was succeeded by the Farm S ...
. These landowners were resettled on more fertile land elsewhere. Today, the South Carolina Sand Hills State Forest comprises about half of the acreage, and half is owned by the
United States Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
as the adjacent Carolina Sandhills National Wildlife Refuge. At first, restoration of forest cover was the goal. Fire suppression was practiced until the 1960s, when prescribed fire was introduced on both the state forest and the Sandhills NWR as part of the restoration of the longleaf/wiregrass ecosystem.
Nokuse Plantation Nokuse Plantation is a nature preserve in northwest Florida's Walton County. It consists of approximately and is the largest privately owned nature preserve in the Southeastern United States. Founded in 2000, it is funded by timber and oil commod ...
is a 53,000-acre private nature preserve located around 100 miles east of
Pensacola, Florida Pensacola () is the westernmost city in the Florida Panhandle, and the county seat and only incorporated city of Escambia County, Florida, United States. As of the 2020 United States census, the population was 54,312. Pensacola is the principal c ...
. The preserve was established by M.C. Davis, a wealthy philanthropist who made his fortune buying and selling land and mineral rights, and who has spent $90 million purchasing land for the preserve, primarily from timber companies. One of its main goals is the restoration of longleaf pine forest, to which end he has had 8 million longleaf pine seedlings planted on the land. A 2009 study by the National Wildlife Federation says that longleaf pine forests will be particularly well adapted to environmental changes caused by climate disruption.


Uses

Vast forests of longleaf pine once were present along the southeastern
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
coast and Gulf Coast of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, as part of the eastern savannas. These forests were the source of naval stores
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on nat ...
,
turpentine Turpentine (which is also called spirit of turpentine, oil of turpentine, terebenthene, terebinthine and (colloquially) turps) is a fluid obtained by the distillation of resin harvested from living trees, mainly pines. Mainly used as a special ...
, and
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
needed by
merchant A merchant is a person who trades in commodities produced by other people, especially one who trades with foreign countries. Historically, a merchant is anyone who is involved in business or trade. Merchants have operated for as long as indust ...
s and the
navy A navy, naval force, or maritime force is the branch of a nation's armed forces principally designated for naval and amphibious warfare; namely, lake-borne, riverine, littoral, or ocean-borne combat operations and related functions. It in ...
for their
ship A ship is a large watercraft that travels the world's oceans and other sufficiently deep waterways, carrying cargo or passengers, or in support of specialized missions, such as defense, research, and fishing. Ships are generally distinguished ...
s. They have been cutover since for timber and usually replaced with faster-growing loblolly pine and slash pine, for
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people ...
, and for urban and suburban development. Due to this deforestation and overharvesting, only about 3% of the original longleaf pine forest remains, and little new is planted. Longleaf pine is available, however, at many nurseries within its range; the southernmost known point of sale is in Lake Worth Beach, Florida. The yellow, resinous
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
is used for
lumber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
and pulp. Boards cut years ago from virgin
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
were very wide, up to , and a thriving salvage business obtains these boards from demolition projects to be reused as flooring in upscale homes. The extremely long needles are popular for use in the ancient craft of coiled basket making. Annual sales of pine straw for use as mulch were estimated at $200M in 2021. The stumps and taproots of old trees become saturated with resin and will not rot. Farmers sometimes find old buried stumps in fields, even in some that were cleared a century ago, and these usually are dug up and sold as
fatwood Fatwood, also known as "fat lighter", "lighter wood", "rich lighter", "pine knot", "lighter knot", "heart pine", "fat stick" or "lighter'd", is derived from the heartwood of pine trees. The stump (and tap root) that is left in the ground aft ...
, "fat lighter", or "lighter wood", which is in demand as kindling for fireplaces, wood stoves, and barbecue pits. In old-growth pine, the heartwood of the bole is often saturated in the same way. When boards are cut from the fat lighter wood, they are very heavy and will not rot, but buildings constructed of them are quite flammable and make extremely hot fires. The seeds of the longleaf pine are edible raw or roasted.


Culture

The Longleaf pine is the official
state tree This is a list of U.S. state, federal district, and territory trees, including official trees of the following of the states, of the federal district, and of the territories. Table See also * List of U.S. state, district, and territorial insign ...
of Alabama. It is commonly but incorrectly believed to be the official state tree of North Carolina, probably because it is referenced by name in the first line of the official
North Carolina State Toast "A Toast" was adopted by the North Carolina General Assembly in 1957: Common uses The State Toast is used at some functions within the University of North Carolina. The University of North Carolina at Chapel Hill's oldest student organization, ...
. Also, the state's highest honor is named the " Order of the Long Leaf Pine". However, the state tree of North Carolina is officially designated as simply "pine," not any particular species of pine.


See also

* Longleaf pine ecosystem * Mountain Longleaf National Wildlife Refuge *
Sonderegger pine Sonderegger is a surname. Notable people with the name include: *Emil Sonderegger Emil Sonderegger (born 28 November 1868 in Herisau – died 15 July 1934) was a Swiss military officer who later became involved in the country's far right pol ...
, a hybrid between loblolly and longleaf species


References


Further reading

* * * * * * * * * * * * * *Cross City: Turpentine Camp—Visit to Aycock & Lindsey Turpentine Camp, Cross City, Florida. By Zora Neale Hurston. August 1939.https://www.floridamemory.com/learn/classroom/learning-units/zora-neale-hurston/documents/essay/


External links

* {{DEFAULTSORT:Pinus Palustris palustris Trees of the Southeastern United States Trees of the South-Central United States Vulnerable plants Trees of the United States Symbols of Alabama Symbols of North Carolina Taxa named by Philip Miller