Biological organisation is the

hierarchy A hierarchy (from Greek: , from , 'president of sacred rites') is an arrangement of items (objects, names, values, categories, etc.) that are represented as being "above", "below", or "at the same level as" one another. Hierarchy is an important ...

of complex

biological Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary in ...

structures and

systems A system is a group of interacting or interrelated elements that act according to a set of rules to form a unified whole. A system, surrounded and influenced by its environment, is described by its boundaries, structure and purpose and express ...

that define life using a reductionistic approach. The traditional hierarchy, as detailed below, extends from atoms to biospheres. The higher levels of this scheme are often referred to as an ecological organisation concept, or as the field, hierarchical ecology. Each level in the hierarchy represents an increase in organisational

complexity Complexity characterises the behaviour of a system or model whose components interaction, interact in multiple ways and follow local rules, leading to nonlinearity, randomness, collective dynamics, hierarchy, and emergence. The term is generall ...

, with each "object" being primarily composed of the previous level's basic unit. The basic principle behind the organisation is the concept of ''

emergence In philosophy, systems theory, science, and art, emergence occurs when an entity is observed to have properties its parts do not have on their own, properties or behaviors that emerge only when the parts interact in a wider whole. Emergence ...

''—the properties and functions found at a hierarchical level are not present and irrelevant at the lower levels. The biological organisation of life is a fundamental premise for numerous areas of scientific research, particularly in the

medical sciences Medicine is the science and practice of caring for a patient, managing the diagnosis, prognosis, prevention, treatment, palliation of their injury or disease, and promoting their health. Medicine encompasses a variety of health care practice ...

. Without this necessary degree of organisation, it would be much more difficult—and likely impossible—to apply the study of the effects of various

physical Physical may refer to: *Physical examination In a physical examination, medical examination, or clinical examination, a medical practitioner examines a patient for any possible medical signs or symptoms of a medical condition. It generally co ...

and chemical phenomena to

diseases A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that ar ...

and physiology (body function). For example, fields such as

cognitive Cognition refers to "the mental action or process of acquiring knowledge and understanding through thought, experience, and the senses". It encompasses all aspects of intellectual functions and processes such as: perception, attention, thought, ...

and behavioral neuroscience could not exist if the brain was not composed of specific types of cells, and the basic concepts of

pharmacology Pharmacology is a branch of medicine, biology and pharmaceutical sciences concerned with drug or medication action, where a drug may be defined as any artificial, natural, or endogenous (from within the body) molecule which exerts a biochemica ...

could not exist if it was not known that a change at the cellular level can affect an entire organism. These applications extend into the ecological levels as well. For example, DDT's direct insecticidal effect occurs at the

subcellular The cell is the basic structural and functional unit of life forms. Every cell consists of a cytoplasm enclosed within a membrane, and contains many biomolecules such as proteins, DNA and RNA, as well as many small molecules of nutrients and ...

level, but affects higher levels up to and including multiple ecosystems. Theoretically, a change in one atom could change the entire biosphere.

Levels

The simple standard biological organisation scheme, from the lowest level to the highest level, is as follows: More complex schemes incorporate many more levels. For example, a molecule can be viewed as a grouping of

elements Element or elements may refer to: Science * Chemical element, a pure substance of one type of atom * Heating element, a device that generates heat by electrical resistance * Orbital elements, parameters required to identify a specific orbit of ...

, and an atom can be further divided into

subatomic particle In physical sciences, a subatomic particle is a particle that composes an atom. According to the Standard Model of particle physics, a subatomic particle can be either a composite particle, which is composed of other particles (for example, a pr ...

s (these levels are outside the scope of biological organisation). Each level can also be broken down into its own hierarchy, and specific types of these biological objects can have their own hierarchical scheme. For example, genomes can be further subdivided into a hierarchy of genes. Each level in the hierarchy can be described by its lower levels. For example, the organism may be described at any of its component levels, including the atomic, molecular, cellular, histological (tissue), organ and organ system levels. Furthermore, at every level of the hierarchy, new functions necessary for the control of life appear. These new roles are not functions that the lower level components are capable of and are thus referred to as ''

emergent properties In philosophy, systems theory, science, and art, emergence occurs when an entity is observed to have properties its parts do not have on their own, properties or behaviors that emerge only when the parts interact in a wider whole. Emergence ...

''. Every organism is organised, though not necessarily to the same degree. An organism can not be organised at the histological (tissue) level if it is not composed of tissues in the first place.

Emergence of biological organisation

Biological organisation is thought to have emerged in the early RNA world when

RNA Ribonucleic acid (RNA) is a polymeric molecule essential in various biological roles in coding, decoding, regulation and expression of genes. RNA and deoxyribonucleic acid ( DNA) are nucleic acids. Along with lipids, proteins, and carbohydra ...

chains began to express the basic conditions necessary for natural selection to operate as conceived by

Darwin Darwin may refer to: Common meanings * Charles Darwin (1809–1882), English naturalist and writer, best known as the originator of the theory of biological evolution by natural selection * Darwin, Northern Territory, a territorial capital city i ...

: heritability, variation of type, and competition for limited resources. Fitness of an RNA replicator (its per capita rate of increase) would likely have been a function of adaptive capacities that were intrinsic (in the sense that they were determined by the nucleotide sequence) and the availability of resources.Michod RE. (2000) Darwinian Dynamics: Evolutionary Transitions in Fitness and Individuality. Princeton University Press, Princeton, New Jersey The three primary adaptive capacities may have been (1) the capacity to replicate with moderate fidelity (giving rise to both heritability and variation of type); (2) the capacity to avoid decay; and (3) the capacity to acquire and process resources. These capacities would have been determined initially by the folded configurations of the RNA replicators (see “ Ribozyme”) that, in turn, would be encoded in their individual nucleotide sequences. Competitive success among different RNA replicators would have depended on the relative values of these adaptive capacities. Subsequently, among more recent organisms competitive success at successive levels of biological organisation, presumably continued to depend, in a broad sense, on the relative values of these adaptive capacities.

Fundamentals

Empirically, a large proportion of the (complex) biological systems we observe in nature exhibit hierarchical structure. On theoretical grounds we could expect complex systems to be hierarchies in a world in which complexity had to evolve from simplicity.

System A system is a group of Interaction, interacting or interrelated elements that act according to a set of rules to form a unified whole. A system, surrounded and influenced by its environment (systems), environment, is described by its boundaries, ...

hierarchies analysis performed in the 1950s, laid the empirical foundations for a field that would be, from the 1980s, hierarchical ecology. The theoretical foundations are summarized by thermodynamics. When

biological systems A biological system is a complex network which connects several biologically relevant entities. Biological organization spans several scales and are determined based different structures depending on what the system is. Examples of biological syst ...

are modeled as

physical system A physical system is a collection of physical objects. In physics, it is a portion of the physical universe chosen for analysis. Everything outside the system is known as the environment. The environment is ignored except for its effects on the ...

s, in its most general abstraction, they are thermodynamic open systems that exhibit

self-organised Self-organization, also called spontaneous order in the social sciences, is a process where some form of overall order arises from local interactions between parts of an initially disordered system. The process can be spontaneous when suffic ...

behavior, and the set/subset relations between

dissipative structures A dissipative system is a thermodynamically open system which is operating out of, and often far from, thermodynamic equilibrium in an environment with which it exchanges energy and matter. A tornado may be thought of as a dissipative system. Dis ...

can be characterized in a hierarchy. A simpler and more direct way to explain the fundamentals of the "hierarchical organisation of life", was introduced in Ecology by Odum and others as the " Simon's hierarchical principle"; SimonSimon's texts at
polaris.gseis.ucla.edu/pagre/simon
o
johncarlosbaez/2011/08/29 transcriptions
emphasized that hierarchy "''emerges almost inevitably through a wide variety of evolutionary processes, for the simple reason that hierarchical structures are stable''". To motivate this deep idea, he offered his "parable" about imaginary watchmakers. :

Notes

References

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