''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

in Equisetaceae, a

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

of ferns, which reproduce by

spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, ...

s rather than seeds. ''Equisetum'' is a "

living fossil A living fossil is an extant taxon that cosmetically resembles related species known only from the fossil record. To be considered a living fossil, the fossil species must be old relative to the time of origin of the extant clade. Living foss ...

", the only living genus of the entire subclass

Equisetidae Equisetidae is one of the four subclasses of Polypodiopsida (ferns), a group of vascular plants with a fossil record going back to the Devonian. They are commonly known as horsetails. They typically grow in wet areas, with whorls of needle-lik ...

, which for over 100 million years was much more diverse and dominated the understorey of late

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ...

forests. Some equisetids were large

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s reaching to tall. The genus ''

Calamites ''Calamites'' is a genus of extinct arborescent (tree-like) horsetails to which the modern horsetails (genus '' Equisetum'') are closely related. Unlike their herbaceous modern cousins, these plants were medium-sized trees, growing to heights o ...

'' of the family

Calamitaceae Calamitaceae is an extinct family of plants related to the modern horsetails. Some members of this family attained tree-like stature during the Carboniferous Period (around ) and in Permian Period, reaching heights of up to 20 meters. The famil ...

, for example, is abundant in

coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal is formed when ...

deposits from the

Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferou ...

period. The pattern of spacing of nodes in horsetails, wherein those toward the apex of the shoot are increasingly close together, is said to have inspired John Napier to invent

logarithm In mathematics, the logarithm is the inverse function to exponentiation. That means the logarithm of a number to the base is the exponent to which must be raised, to produce . For example, since , the ''logarithm base'' 10 ...

s. Modern horsetails first appeared during the

Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...

period. A superficially similar but entirely unrelated

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...

genus, mare's tail (''

Hippuris ''Hippuris'', the mare's tail, was previously the sole genus in the family Hippuridaceae. Following genetic research by the Angiosperm Phylogeny Group, it has now been transferred to the family Plantaginaceae, with Hippuridaceae being reduced t ...

''), is occasionally referred to as "horsetail", and adding to confusion, the name "mare's tail" is sometimes applied to ''Equisetum''. Despite centuries of use in

traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...

, there is no evidence that ''Equisetum'' has any

medicinal properties Evidence-based medicine (EBM) is "the conscientious, explicit and judicious use of current best evidence in making decisions about the care of individual patients". The aim of EBM is to integrate the experience of the clinician, the values of t ...

Etymology

The name "horsetail", often used for the entire group, arose because the branched species somewhat resemble a

horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...

's tail. Similarly, the

scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...

''Equisetum'' is derived from the

Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...

('horse') + ('bristle'). Other names include candock for branching species, and snake grass or scouring-rush for unbranched or sparsely branched species. The latter name refers to the rush-like appearance of the plants and to the fact that the stems are coated with abrasive

silicate In chemistry, a silicate is any member of a family of polyatomic anions consisting of silicon and oxygen, usually with the general formula , where . The family includes orthosilicate (), metasilicate (), and pyrosilicate (, ). The name is a ...

s, making them useful for scouring (cleaning) metal items such as cooking pots or drinking mugs, particularly those made of tin. '' E. hyemale'', rough horsetail, is still boiled and then dried in

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...

to be used for the final polishing process on

woodcraft The term woodcraft — or woodlore — denotes bushcraft skills and experience in matters relating to living and thriving in the woods—such as hunting, fishing, and camping—whether on a short- or long-term basis. Traditionally, woodcraft p ...

to produce a smooth finish. In

German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...

, the corresponding name is ('tin-herb'). In Spanish-speaking countries, these plants are known as ('horsetail').

Description

''Equisetum''

leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

are greatly reduced and usually non-

photosynthetic Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...

. They contain a single, non-branching vascular trace, which is the defining feature of

microphyll In plant anatomy and evolution a microphyll (or lycophyll) is a type of plant leaf with one single, unbranched leaf vein. Plants with microphyll leaves occur early in the fossil record, and few such plants exist today. In the classical concept of ...

s. However, it has recently been recognised that horsetail microphylls are probably not ancestral as in lycophytes (clubmosses and relatives), but rather derived

adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...

s, evolved by reduction of megaphylls. The leaves of horsetails are arranged in whorls fused into nodal sheaths. The stems are usually green and photosynthetic, and are distinctive in being hollow, jointed and ridged (with sometimes 3 but usually 6–40 ridges). There may or may not be whorls of branches at the nodes. Unusually, the branches often emerge below the leaves in an internode, and grow from buds between their bases.

Spores

The

s are borne under

sporangiophore {{Short pages monitor