family (biology)
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Family (, : ) is one of the eight major hierarchical
taxonomic rank In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of or ...
s in Linnaean taxonomy. It is classified between order and
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community for extended periods. The continual publication of new data and diverse opinions plays a crucial role in facilitating adjustments and ultimately reaching a consensus over time.


Nomenclature

The naming of families is codified by various international bodies using the following suffixes: *In fungal, algal, and botanical nomenclature, the family names of plants, fungi, and algae end with the suffix " -aceae", except for a small number of historic but widely used names including Compositae and Gramineae. *In zoological nomenclature, the family names of animals end with the suffix " -idae".
/ref> Name changes at the family level are regulated by the codes of nomenclature. For botanical families, some traditional names like Palmae ( Arecaceae), Cruciferae ( Brassicaceae), and Leguminosae ( Fabaceae) are conserved alongside their standardized -aceae forms due to their historical significance and widespread use in the literature. Family names are typically formed from the stem of a type genus within the family. In zoology, when a valid family name is based on a genus that is later found to be a junior synonym, the family name may be maintained for stability if it was established before 1960. In botany, some family names that were found to be junior synonyms have been conserved due to their widespread use in the scientific literature. The family-group in zoological nomenclature includes several ranks: superfamily (-oidea), family (-idae),
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
(-inae), and tribe (-ini). Under the principle of coordination, a name established at any of these ranks can be moved to another rank while retaining its original authorship and date, requiring only a change in suffix to reflect its new rank. New family descriptions are relatively rare in taxonomy, occurring in fewer than one in a hundred taxonomic publications. Such descriptions typically result from either the discovery of organisms with unique combinations of characters that do not fit existing families, or from phylogenetic analyses that reveal the need for reclassification.


History

The taxonomic term ' was first used by French botanist Pierre Magnol in his ' (1689) where he called the seventy-six groups of plants he recognised in his tables families ('). The concept of rank at that time was not yet settled, and in the preface to the ' Magnol spoke of uniting his families into larger ', which is far from how the term is used today. In his work ''Philosophia Botanica'' published in 1751,
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
employed the term ''familia'' to categorize significant plant groups such as trees, herbs, ferns, palms, and so on. Notably, he restricted the use of this term solely within the book's morphological section, where he delved into discussions regarding the vegetative and generative aspects of plants. Subsequently, in French botanical publications, from Michel Adanson's ' (1763) and until the end of the 19th century, the word ' was used as a French equivalent of the Latin ' (or '). The family concept in botany was further developed by the French botanists Antoine Laurent de Jussieu and Michel Adanson. Jussieu's 1789 ''Genera Plantarum'' divided plants into 100 'natural orders,' many of which correspond to modern plant families. However, the term 'family' did not become standardized in botanical usage until after the mid-nineteenth century. In
zoology Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...
, the family as a rank intermediate between order and genus was introduced by
Pierre André Latreille Pierre André Latreille (; 29 November 1762 – 6 February 1833) was a French zoology, zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained hi ...
in his ' (1796). He used families (some of them were not named) in some but not in all his orders of "insects" (which then included all
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s). The standardization of zoological family names began in the early nineteenth century. A significant development came in 1813 when William Kirby introduced the -idae suffix for animal family names, derived from the Greek 'eidos' meaning 'resemblance' or 'like'. The adoption of this naming convention helped establish families as an important taxonomic rank. By the mid-1800s, many of Linnaeus's broad genera were being elevated to family status to accommodate the rapidly growing number of newly discovered species. In nineteenth-century works such as the ' of Augustin Pyramus de Candolle and the ' of George Bentham and Joseph Dalton Hooker this word ' was used for what now is given the rank of family.


Uses

Families serve as valuable units for evolutionary, paleontological, and genetic studies due to their relatively greater stability compared to lower taxonomic levels like genera and species. Families play a significant practical role in biological education and research. They provide an efficient framework for teaching taxonomy, as they group organisms with general similarities while remaining specific enough to be useful for identification purposes. For example, in botany, learning the characteristics of major plant families helps students identify related species across different geographic regions, since families often have worldwide distribution patterns. In many groups of organisms, families serve as the primary level for taxonomic identification keys, making them particularly valuable for field guides and systematic work as they often represent readily recognizable groups of related organisms with shared characteristics. In ecological and biodiversity research, families frequently serve as the foundational level for identification in survey work and environmental studies. This is particularly useful because families often share life history traits or occupy similar ecological niches. Some families show strong correlations between their taxonomic grouping and ecological functions, though this relationship varies among different groups of organisms. The stability of family names has practical importance for applied biological work, though this stability faces ongoing challenges from new scientific findings. Modern molecular studies and phylogenetic analyses continue to refine the understanding of family relationships, sometimes leading to reclassification. The impact of these changes varies among different groups of organisms – while some families remain well-defined and easily recognizable, others require revision as new evidence emerges about evolutionary relationships. This balance between maintaining nomenclatural stability and incorporating new scientific discoveries remains an active area of taxonomic practice.


See also

*
Systematics Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...
, the study of the diversity of living organisms * Cladistics, the classification of organisms by their order of branching in an evolutionary tree * Virus classification * List of Anuran families * List of Testudines families * List of fish families * List of families of spiders


References


Bibliography

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External links

* {{DEFAULTSORT:rank13