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Dog anatomy comprises the anatomical studies of the visible parts of the body of a
domestic dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is Domestication of the dog, derived from the extinct Pleistocene wolf, and the modern wolf is the dog's n ...
. Details of structures vary tremendously from
breed A breed is a specific group of domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist several slig ...
to breed, more than in any other animal species, wild or domesticated, as dogs are highly variable in height and weight. The smallest known adult dog was a Yorkshire Terrier that stood only at the shoulder, in length along the head and body, and weighed only . The heaviest dog was an
English Mastiff The English Mastiff, or simply the Mastiff, is a British dog breed of very large size. Likely descended from the ancient Alaunt and Pugnaces Britanniae, with a significant input from the Alpine Mastiff in the 19th century. Distinguished by i ...
named Zorba which weighed . The tallest known adult dog is a
Great Dane The Great Dane is a large sized dog breed originating from Germany. The Great Dane descends from hunting dogs from the Middle Ages used to hunt wild boar and deer, and as guardians of German nobility. It is one of the largest breeds in the worl ...
that stands at the shoulder.


Anatomy

Source:


Muscles

The following is a list of the muscles in the dog, along with their origin, insertion, action and innervation. ''Extrinsic muscles of the thoracic limb and related structures:'' Descending superficial pectoral: originates on the first sternebrae and inserts on the greater tubercle of the humerus. It both adducts the limb and also prevents the limb from being abducted during weight bearing. It is innervated by the cranial pectoral nerves. Transverse superficial pectoral: originates on the second and third sternebrae and inserts on the greater tubercle of the humerus. It also adducts the limb and prevents the limb from being abducted during weight bearing. It is innervated by the cranial pectoral nerves. Deep pectoral: originates on the ventral sternum and inserts on the lesser tubercle of the humerus. It acts to extend the shoulder joint during weight bearing and flexes the shoulder when there is no weight. It is innervated by the caudal pectoral nerves. Sternocephalicus: originates on the sternum and inserts on the temporal bone of the head. Its function is to move the head and neck from side to side. It is innervated by the accessory nerve. Sternohyoideus: originates on the sternum and inserts on the basihyoid bone. Its function is to move the tongue caudally. It is innervated by the ventral branches of the cervical spinal nerves. Sternothyoideus: originates on the first coastal cartilage and inserts on the thyroid cartilage. Its function is also to move the tongue caudally. It is innervated by the ventral branches of the cervical spinal nerves. Omotransversarius: originates on the spine of the scapula and inserts on the wing of the atlas. Its function is to advance the limb and flex the neck laterally. It is innervated by the accessory nerve. Trapezius: originates on the supraspinous ligament and inserts on the spine of the scapula. Its function is to elevate and abduct the forelimb. It is innervated by the accessory nerve. Rhomboideus: originates on the nuchal crest of the occipital bone and inserts on the scapula. Its function is to elevate the forelimb. It is innervated by the ventral branches of the spinal nerves. Latissimus dorsi: originates on thoracolumbar fascia and inserts on the teres major tuberosity of the humerus. Its function is to flex the shoulder joint. It is innervated by the thoracodorsal nerve. Serratus ventralis: originates on the transverse processes of the last 5 cervical vertebrae and inserts on the scapula. Its function is to support the trunk and depress the scapula. It is innervated by the ventral branches of the cervical spinal nerves. ''Intrinsic muscles of the thoracic limb:'' Deltoideus: originates on the acromial process of the scapula and inserts on the deltoid tuberosity. It acts to flex the shoulder. It is innervated by the axillary nerve. Infraspinatus: originates on the infraspinatus fossa and inserts on the greater tubercle of the humerus. It acts to extend and flex the shoulder joint. It is innervated by the suprascapular nerve. Teres minor: originates on the infra glenoid tubercle on the scapula and inserts on the teres minor tuberosity of the humerus. It acts to flex the shoulder and rotate the arm laterally. It is innervated by the axillary nerve. Supraspinatus: originates on the supraspinous fossa and inserts on the greater tubercle of the humerus. It acts to extend and stabilize the shoulder joint. It is innervated by the suprascapular nerve. ''Medial muscles of the scapula and shoulder:'' Subscapularis: originates on the subscapular fossa and inserts on the greater tubercle of the humerus. It acts to rotate the arm medially and stabilize the joint. It is innervated by the subscapular nerve. Teres major: originates on the scapula and inserts on the teres major tuberosity of the humerus. It acts to flex the shoulder and rotate the arm medially. It is innervated by the axillary nerve. Coracobrachialis: originates on the coracoid process of the scapula and inserts on the crest of the lesser tubercle of the humerus. It acts to adduct, extend and stabilize the shoulder joint. It is innervated by the musculocutaneous nerve. ''Caudal muscles of brachium:'' Tensor fasciae antebrachium: originates on the fascia covering the latissimus dorsi and inserts on the olecranon. It acts to extend the elbow. It is innervated by the radial nerve. Triceps brachii: originates on the caudal border of the scapula and inserts on the olecranon tuber. It acts to extend the elbow and flex the shoulder. It is innervated by the radial nerve. Anconeus: originates on the humerus and inserts on the proximal end of the ulna. It acts to extend the elbow. It is innervated by the radial nerve. ''Cranial muscles of the arm:'' Biceps brachia: originates on the supraglenoid tubercle and inserts on the ulnar and radial tuberosities. It acts to flex the elbow and extend the shoulder. It is innervated by the musculocutaneous nerve. Brachialis: originates on the lateral surface of humerus and inserts on the ulnar and radial tuberosities. It acts to flex the elbow. It is innervated by the musculocutaneous nerve. ''Cranial and lateral muscles of antebrachium:'' Extensor carpi radial: originates on the supracondylar crest and inserts on the metacarpals. It acts to extend the carpus. It is innervated by the radial nerve. Common digital extensor: originates on the lateral epicondyle of the humerus and inserts on the distal phalanges. It acts to extend the carpus and joints of the digits 3, 4, and 5. It is innervated by the radial nerve. Extensor carpi ulnar: originates on the lateral epicondyle of the humerus and inserts on the metacarpal 5 and the accessory carpal bone. It acts to abduct and extend the carpal joint. It is innervated by the radial nerve. Supinator: originates on the lateral epicondyle of the humerus and inserts on the radius. It acts to rotate the forearm laterally. It is innervated by the radial nerve. Abductor pollicis longus: originates on the ulna and inserts on metacarpal 1. It acts to abduct the digit and extend the carpal joints. It is innervated by the radial nerve. ''Caudal and medial muscles of forearm:'' Pronator teres: originates on the medial epicondyle of the humerus and inserts on the medial border of the radius. It acts to rotate forearm medially and flex the elbow. It is innervated by the median nerve. Flexor carpi radial: originates on the medial epicondyle of the humerus and inserts on the palmar side of metacarpals 2 and 3. It acts to flex the carpus. It is innervated by the median nerve. Superficial digital flexor: originates on the medial epicondyle of the humerus and inserts on the palmar surface of the middle phalanges. It acts to flex the carpus, metacarpophalangeal and proximal interphalangeal joints of the digits. It is innervated by the median nerve. Flexor carpi ulnar: originates on the olecranon and inserts on the accessory carpal bone. It acts to flex the carpus. It is innervated by the ulnar nerve. Deep digital flexor: originates on the medial epicondyle of the humerus and inserts on the palmar surface of the distal phalanx. It acts to flex the carpus, metacarpophalangeal joints, and the proximal and distal interphalangeal joints of the digits. It is innervated by the median nerve. Pronator quadratus: originates on surfaces of the radius and ulna. It acts to pronate the paw. It is innervated by the median nerve. ''Caudal muscles of the thigh:'' Biceps femoris: originates on the ischiatic tuberosity and inserts on the patellar ligament. It acts to extend the hip, stifle and hock. It is innervated by the sciatic nerve. Semitendinosus: originates on the ischiatic tuberosity and inserts on the tibia. It acts to extend the hip, flex the stifle and extend the hock. It is innervated by the sciatic nerve. Semimembranosus: originates on the ischiatic tuberosity and inserts on the femur and tibia. It acts to extend the hip and stifle. It is innervated by the sciatic nerve. ''Medial muscles of the thigh:'' Sartorius: originates on the ilium and inserts on the patella and tibia. It acts to flex the hip and both flex and extend the stifle. It is innervated by the femoral nerve. Gracilis: originates on the pelvic symphysis and inserts on the cranial border of the tibia. It acts to adduct the limb, flex the stifle and extend the hip and hock. It is innervated by the obturator nerve. Pectineus: originates on the iliopubic eminence and inserts on the caudal femur. It acts to adduct the limb. It is innervated by the obturator nerve. Adductor: originates on the pelvic symphysis and inserts on the lateral femur. It acts to adduct the limb and extend the hip. It is innervated by the obturator nerve. ''Lateral muscles of the pelvis:'' Tensor fasciae latae: originates on the tuber coxae of the ilium and inserts on the lateral femoral fascia. It acts to flex the hip and extend the stifle. It is innervated by the cranial gluteal nerve. Superficial gluteal: originates on the lateral border of the sacrum and inserts on the 3rd trochanter. It acts to extend the hip and abduct the limb. It is innervated by the caudal gluteal nerve. Middle gluteal: originates on the ilium and inserts on the greater trochanter. It acts to abduct the hip and rotate the pelvic limb medially. It is innervated by the cranial gluteal nerve. Deep gluteal: originates on the ischiatic spine and inserts on the greater trochanter. It acts to extend the hip and rotate the pelvic limb medially. It is innervated by the cranial gluteal nerve. ''Caudal hip muscles:'' Internal obturator: originates on the pelvic symphysis and inserts on the trochanteric fossa of the femur. It acts to rotate the pelvic limb laterally. It is innervated by the sciatic nerve. Gemelli: originates on the lateral surface of the ischium and inserts on the trochanteric fossa. It acts to rotate the pelvic limb laterally. It is innervated by the sciatic nerve. Quadratus femoris: originates on the ischium and inserts on the intertrochanteric crest. It acts to extend the hip and rotate the pelvic limb laterally. External obturator: originates on the pubis and ischium and inserts on the trochanteric fossa. It acts to rotate the pelvic limb laterally. It is innervated by the obturator nerve. ''Cranial muscles of the thigh:'' Quadriceps femoris: originates on the femur and the ilium and inserts on the tibial tuberosity. It acts to extend the stifle and to flex the hip. It is innervated by the femoral nerve. Ilipsoas: originates on the ilium and inserts on the lesser trochanter. It acts to flex the hip. It is innervated by the femoral nerve. ''Craniolateral muscles of the leg:'' Cranial tibial: originates on tibia and inserts on the plantar surfaces of metatarsals 1 and 2. It acts to flex the tarsus and rotates the paw laterally. It is innervated by the peroneal nerve. Long digital extensor: originates from the extensor fossa of the femur and inserts on the extensor processes of the distal phalanges. It acts to extend the digits and flex the tarsus. It is innervated by the peroneal nerve. Peroneus longus: originates on both the tibia and fibula and inserts on the 4th tarsal bone and the plantar aspect of the metatarsals. It acts to flex the tarsus and rotate the paw medially. It is innervated by the peroneal nerve. ''Caudal muscles of the leg:'' Gastrocnemius: originates on the supracondylar tuberosities of the femur and inserts on the tuber calcanei. It acts to extend the tarsus and flex the stifle. It is innervated by the tibial nerve. Superficial digital flexor: originates on the lateral supracondylar tuberosity of the femur and inserts on the tuber calcanei and bases of the middle phalanges. It acts to flex the stifle and extend the tarsus. It is innervated by the tibial nerve. Deep digital flexor: originates on the fibular and inserts on the plantar surface of the distal phalanges. It acts to flex the digits and extend the tarsus. It is innervated by the tibial nerve. Popliteus: originates on the lateral condyle of the femur and inserts on the tibia. It acts to rotate the leg medially. It is innervated by the tibial nerve.


Skeleton

''Bones and their significant points for muscle attachment:'' Scapula: Spine of the Scapula, Supraglenoid Tubercle, Glenoid Cavity, Acromion Process, Supraspinous Fossa, Infraspinous Fossa, Neck, Coracoid, Process, Subscapular Fossa Humerus: Head of Humerus, Greater Tubercle, Lesser Tubercle, Intertubercular Groove, Deltopectoral Crest, Deltoid Tuberosity, Body of the Humerus, Epicondyles (Medial and Lateral), Humeral condyle (Trochlea and Capitulum, Radial and Olecranon Fossae) Ulna and Radius: Olecranon Process, Trochlear Notch, Anconeal Process, Coronoid Processes (Medial and Lateral), Body of Ulna, Head of Radius, Body of Radius, Distal Trochlea, Styiloid Process (Medial and Lateral), Interosseus Space Metacarpals: Carpal Bones (Radial and Ulnar), Accessory Carpal Bone, First, Second, Third, and Fourth Metacarpals, Phalanges, Proximal Base, Body, Head, Ungual crest, Ungual process (Nails), Extensor process, Carpometacarpal Joints, Metacarpophalangeal Joints, Proximalinterphalangeal Joints, Interphalangeal Joints Femur: Head, Ligament of Head, Neck, Greater Trochanter, Lesser Trochanter, Trochanteric Fossa, Acetabulum Fossa (on Hip Bone), Distal Femur, Trochlea (and Ridges), Condyles (Medial/Lateral), Epicondyles (Medial/Lateral), Intercondylar Fossa, Extensor Fossa (Tiny Dent), Infrapatellar Fat Pad, Fabellae (Medial/Lateral) Patella kneecap Tibia and Fibula: Tibial Condyles (Medial/Lateral), Intercondylar Eminences, Extensor Notch (Lateral), Tibial Tuberosity (Cranial), Tibial Cochlea, Medial Malleolus, Lateral Malleolus, Head of Fibula Metatarsals: Talus, Calcaneus, Trochlear Ridges, Central Tarsal Bone, First, Second, and Third Tarsal Bones Vertebra Body, Pedicles, Laminae, Spinous Process, Transverse Process (Wings), Articular Process, Vertebral Foramen, Intervertebral Foramina, Atlas (C1), Axis (C2), dens, Ventral Lamina (on C6) Pelvis: Acetabulum, Ilium, Ischium, Pubis File:Dog anatomy lateral skeleton view.jpg, Lateral view of a dog skeleton File:Skull of a dog.png, Lateral view of a dog skull - jaws open File:Skull of a dog diagram2.svg, Lateral view of a dog skull File:Dog Skull.JPG, Frontal view of a dog skull File:Azawakh K9.jpg, Image of dog teeth File:Dog skull (MAV) 02.jpg, Skull of a dog Skull In 1986, a study of skull morphology found that the domestic dog is morphologically distinct from all other canids except the wolf-like canids. The difference in size and proportion between some breeds are as great as those between any wild genera, but all dogs are clearly members of the same species. In 2010, a study of dog skull shape compared to extant carnivorans proposed that "The greatest shape distances between dog breeds clearly surpass the maximum divergence between species in the Carnivora. Moreover, domestic dogs occupy a range of novel shapes outside the domain of wild carnivorans." The domestic dog compared to the wolf shows the greatest variation in the size and shape of the skull (Evans 1979) that range from 7 to 28 cm in length (McGreevy 2004). Wolves are dolichocephalic (long skulled) but not as extreme as some breeds of dogs such as greyhounds and Russian wolfhounds (McGreevy 2004). Canine
brachycephaly Brachycephaly (derived from the Ancient Greek '' βραχύς'', 'short' and '' κεφαλή'', 'head') is the shape of a skull shorter than typical for its species. It is perceived as a desirable trait in some domesticated dog and cat breeds, ...
(short-skulledness) is found only in domestic dogs and is related to
paedomorphosis Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compare ...
(Goodwin 1997). Puppies are born with short snouts, with the longer skull of dolichocephalic dogs emerging in later development (Coppinger 1995). Other differences in head shape between brachycephalic and dolichocephalic dogs include changes in the craniofacial angle (angle between the basilar axis and
hard palate The hard palate is a thin horizontal bony plate made up of two bones of the facial skeleton, located in the roof of the mouth. The bones are the palatine process of the maxilla and the horizontal plate of palatine bone. The hard palate spans t ...
) (Regodón 1993), morphology of the
temporomandibular joint In anatomy, the temporomandibular joints (TMJ) are the two joints connecting the jawbone to the skull. It is a bilateral synovial articulation between the temporal bone of the skull above and the mandible below; it is from these bones that it ...
(Dickie 2001), and radiographic anatomy of the
cribriform plate In mammalian anatomy, the cribriform plate (Latin for lit. ''sieve-shaped''), horizontal lamina or lamina cribrosa is part of the ethmoid bone. It is received into the ethmoidal notch of the frontal bone and roofs in the nasal cavities. It supp ...
(Schwarz 2000). All cited in Roberts. One study found that the relative reduction in dog skull length compared to its width (the Cephalic Index) was significantly correlated to both the position and the angle of the brain within the skull. This was regardless of the brain size or the body weight of the dog.


Respiratory system

The
respiratory system The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and plants. The anatomy and physiology that make this happen varies g ...
is the set of organs responsible for the intake of oxygen and the expelling of carbon dioxide. As dogs have few
sweat glands Sweat glands, also known as sudoriferous or sudoriparous glands, , are small tubular structures of the skin that produce sweat. Sweat glands are a type of exocrine gland, which are glands that produce and secrete substances onto an epithelial sur ...
in their skin, the respiratory system also plays an important role in body
thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperatur ...
. Dogs are mammals with two large
lungs The lungs are the primary organs of the respiratory system in humans and most other animals, including some snails and a small number of fish. In mammals and most other vertebrates, two lungs are located near the backbone on either si ...
that are further divided into lobes. They have a spongy appearance due to the presence of a system of delicate branches of the
bronchioles The bronchioles or bronchioli (pronounced ''bron-kee-oh-lee'') are the smaller branches of the bronchial airways in the lower respiratory tract. They include the terminal bronchioles, and finally the respiratory bronchioles that mark the start ...
in each lung, ending in closed, thin-walled chambers (the points of gas exchange) called
alveoli Alveolus (; pl. alveoli, adj. alveolar) is a general anatomical term for a concave cavity or pit. Uses in anatomy and zoology * Pulmonary alveolus, an air sac in the lungs ** Alveolar cell or pneumocyte ** Alveolar duct ** Alveolar macrophage * M ...
. The presence of a muscular structure, the
diaphragm Diaphragm may refer to: Anatomy * Thoracic diaphragm, a thin sheet of muscle between the thorax and the abdomen * Pelvic diaphragm or pelvic floor, a pelvic structure * Urogenital diaphragm or triangular ligament, a pelvic structure Other * Diap ...
, exclusive to mammals, divides the
peritoneal cavity The peritoneal cavity is a potential space between the parietal peritoneum (the peritoneum that surrounds the abdominal wall) and visceral peritoneum (the peritoneum that surrounds the internal organs). The parietal and visceral peritonea are la ...
from the
pleural cavity The pleural cavity, pleural space, or interpleural space is the potential space between the pleurae of the pleural sac that surrounds each lung. A small amount of serous pleural fluid is maintained in the pleural cavity to enable lubrication b ...
, besides assisting the lungs during inhalation.


Digestive system

The organs that make up the canine
digestive system The human digestive system consists of the gastrointestinal tract plus the accessory organs of digestion (the tongue, salivary glands, pancreas, liver, and gallbladder). Digestion involves the breakdown of food into smaller and smaller compone ...
are: *
Mouth In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on ...
*
Tongue The tongue is a muscular organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive process, and is the primary organ of taste. The tongue's upper surface (dorsum) is covered by taste ...
*
Esophagus The esophagus (American English) or oesophagus (British English; both ), non-technically known also as the food pipe or gullet, is an organ in vertebrates through which food passes, aided by peristaltic contractions, from the pharynx to t ...
*
Stomach The stomach is a muscular, hollow organ in the gastrointestinal tract of humans and many other animals, including several invertebrates. The stomach has a dilated structure and functions as a vital organ in the digestive system. The stomach i ...
*
Liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
*
Pancreas The pancreas is an organ of the digestive system and endocrine system of vertebrates. In humans, it is located in the abdomen behind the stomach and functions as a gland. The pancreas is a mixed or heterocrine gland, i.e. it has both an ...
*
Large intestine The large intestine, also known as the large bowel, is the last part of the gastrointestinal tract and of the digestive system in tetrapods. Water is absorbed here and the remaining waste material is stored in the rectum as feces before bein ...
*
Small intestine The small intestine or small bowel is an organ (anatomy), organ in the human gastrointestinal tract, gastrointestinal tract where most of the #Absorption, absorption of nutrients from food takes place. It lies between the stomach and large intes ...
*
Rectum The rectum is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. The adult human rectum is about long, and begins at the rectosigmoid junction (the end of the sigmoid colon) at the l ...
*
Anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, ...
File:Dog cecum.jpg, Dog
cecum The cecum or caecum is a pouch within the peritoneum that is considered to be the beginning of the large intestine. It is typically located on the right side of the body (the same side of the body as the appendix, to which it is joined). The wo ...
. File:Dog digestive tract.jpg, Dog
digestive tract The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans and ...
. File:Dog stomach.jpg, Dog
stomach The stomach is a muscular, hollow organ in the gastrointestinal tract of humans and many other animals, including several invertebrates. The stomach has a dilated structure and functions as a vital organ in the digestive system. The stomach i ...
. File:Dog stomach (open, inner view).jpg, Dog stomach (open, inner view). File:Technique of formalin fixation applied to dog tongue.jpg, Technique of formalin fixation applied to dog
tongue The tongue is a muscular organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive process, and is the primary organ of taste. The tongue's upper surface (dorsum) is covered by taste ...
. File:Dog ilium.jpg, Dog
ileum The ileum () is the final section of the small intestine in most higher vertebrates, including mammals, reptiles, and birds. In fish, the divisions of the small intestine are not as clear and the terms posterior intestine or distal intestine m ...
. File:Dog liver.jpg, Vascular structure of the dog
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
.



Physical characteristics

Sixty percent of the dog's body mass falls on the front legs. Like most predatory mammals, the dog has powerful muscles, a
cardiovascular The blood circulatory system is a system of organs that includes the heart, blood vessels, and blood which is circulated throughout the entire body of a human or other vertebrate. It includes the cardiovascular system, or vascular system, tha ...
system that supports both sprinting and endurance and teeth for catching, holding, and tearing. The dog's muscles provides the ability to jump and leap. Their legs can propel them forward rapidly, leaping as necessary to chase and overcome prey. They have small, tight feet, walking on their toes (thus having a
digitigrade In terrestrial vertebrates, digitigrade () locomotion is walking or running on the toes (from the Latin ''digitus'', 'finger', and ''gradior'', 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the groun ...
stance and locomotion). Their rear legs are fairly rigid and sturdy. The front legs are loose and flexible with only muscle attaching them to the torso. The dog's muzzle size will vary with the breed. Dogs with medium muzzles, such as the German Shepherd Dog, are called mesocephalic and dogs with a pushed in muzzle, such as the Pug, are called brachycephalic. Today's
toy A toy or plaything is an object that is used primarily to provide entertainment. Simple examples include toy blocks, board games, and dolls. Toys are often designed for use by children, although many are designed specifically for adults and pet ...
breeds have skeletons that mature in only a few months, while giant breeds, such as the
Mastiff A mastiff is a large and powerful type of dog. Mastiffs are among the largest dogs, and typically have a short coat, a long low-set tail and large feet; the skull is large and bulky, the muzzle broad and short ( brachycephalic) and the ears dr ...
s, take 16 to 18  months for the skeleton to mature.
Dwarfism Dwarfism is a condition wherein an organism is exceptionally small, and mostly occurs in the animal kingdom. In humans, it is sometimes defined as an adult height of less than , regardless of sex; the average adult height among people with dw ...
has affected the proportions of some breeds' skeletons, as in the
Basset Hound The Basset Hound is a short-legged breed of dog in the hound family. The Basset is a scent hound that was originally bred for the purpose of hunting hare. Their sense of smell and ability to ''ground-scent'' is second only to the Bloodhound.Har ...
. All dogs (and all living
Canidae Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...
) have a ligament connecting the spinous process of their first thoracic (or chest) vertebra to the back of the axis bone (second cervical or neck bone), which supports the weight of the head without active muscle exertion, thus saving energy.Wang, Xiaoming and Tedford, Richard H. Dogs: Their Fossil Relatives and Evolutionary History. New York: Columbia University Press, 2008. pp.97-8 This ligament is analogous in function (but different in exact structural detail) to the
nuchal ligament The nuchal ligament is a ligament at the back of the neck that is continuous with the supraspinous ligament. Structure The nuchal ligament extends from the external occipital protuberance on the skull and median nuchal line to the spinous proc ...
found in
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...
s. This ligament allows dogs to carry their heads while running long distances, such as while following scent trails with their nose to the ground, without expending much energy. Dogs have disconnected shoulder bones (lacking the collar bone of the human skeleton) that allow a greater stride length for running and leaping. They walk on four toes, front and back, and have vestigial
dewclaw A dewclaw is a digit – vestigial in some animals – on the foot of many mammals, birds, and reptiles (including some extinct orders, like certain theropods). It commonly grows higher on the leg than the rest of the foot, such that in digit ...
s on their front legs and on their rear legs. When a dog has extra dewclaws in addition to the usual one in the rear, the dog is said to be "double dewclawed."


Size

Dogs are highly variable in height and weight. The smallest known adult dog was a Yorkshire Terrier that stood only at the shoulder, in length along the head and body, and weighed only . The largest known adult dog was an
English Mastiff The English Mastiff, or simply the Mastiff, is a British dog breed of very large size. Likely descended from the ancient Alaunt and Pugnaces Britanniae, with a significant input from the Alpine Mastiff in the 19th century. Distinguished by i ...
which weighed . The tallest known adult dog is a
Great Dane The Great Dane is a large sized dog breed originating from Germany. The Great Dane descends from hunting dogs from the Middle Ages used to hunt wild boar and deer, and as guardians of German nobility. It is one of the largest breeds in the worl ...
that stands at the shoulder. In 2007, a study identified a
gene In biology, the word gene (from , ; "...Wilhelm Johannsen coined the word gene to describe the Mendelian units of heredity..." meaning ''generation'' or ''birth'' or ''gender'') can have several different meanings. The Mendelian gene is a b ...
that is proposed as being responsible for size. The study found a
regulatory sequence A regulatory sequence is a segment of a nucleic acid molecule which is capable of increasing or decreasing the expression of specific genes within an organism. Regulation of gene expression is an essential feature of all living organisms and ...
next to the gene
Insulin-like growth factor 1 Insulin-like growth factor 1 (IGF-1), also called somatomedin C, is a hormone similar in molecular structure to insulin which plays an important role in childhood growth, and has anabolic effects in adults. IGF-1 is a protein that in humans ...
(IGF1) and together with the gene and regulatory sequence "is a major contributor to body size in all small dogs." Two variants of this gene were found in large dogs, making a more complex reason for large breed size. The researchers concluded this gene's instructions to make dogs small must be at least 12,000  years old and it is not found in wolves. Another study has proposed that
lap dog A lap dog or lapdog is a dog that is both small enough to be held in the arms or lie comfortably on a person's lap and temperamentally predisposed to doing so. ''Lapdog'' is not a specific breed, but a generic term for a type of dog that is sm ...
s (small dogs) are among the oldest existing
dog type Dog types are broad categories of domestic dogs based on form, function, or style of work, lineage, or appearance. Some may be locally adapted dog types (or '' landraces'') that may have the visual characteristics of a modern purebred dog. In ...
s.


Coat

Domestic dogs often display the remnants of countershading, a common natural camouflage pattern. The general theory of countershading is that an animal that is lit from above will appear lighter on its upper half and darker on its lower half where it will usually be in its own shade. This is a pattern that predators can learn to watch for. A counter shaded animal will have dark coloring on its upper surfaces and light coloring below. This reduces the general visibility of the animal. One reminder of this pattern is that many breeds will have the occasional "blaze", stripe, or "star" of white fur on their chest or undersides. A study found that the genetic basis that explains coat colors in horse coats and cat coats did not apply to dog coats. The project took samples from 38 different breeds to find the gene (a
beta defensin Beta defensins are a family of vertebrate defensins. The beta defensins are antimicrobial peptides implicated in the resistance of epithelial surfaces to microbial colonization. Defensins are 2-6 kDa, cationic, microbicidal peptides active agai ...
gene) responsible for dog coat color. One version produces yellow dogs and a mutation produces black. All dog coat colors are modifications of black or yellow. For example, the white in white
miniature schnauzer The Miniature Schnauzer is a breed of small dog of the Schnauzer type that originated in Germany in the mid-to-late 19th century. Miniature Schnauzers may have been developed from the smallest specimens of the Standard Schnauzer, or crosses be ...
s is a cream color, not
albinism Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino. Varied use and interpretation of the term ...
(a genotype of e/e at MC1R.) Modern dog breeds exhibit a diverse array of fur coats, including dogs without fur, such as the
Mexican Hairless Dog The Xoloitzcuintle (or Xoloitzquintle, Xoloitzcuintli, or Xolo) is one of several breeds of hairless dog. It is found in standard, intermediate, and miniature sizes. The Xolo also comes in a coated variety, totally covered in fur. Coated and hair ...
. Dog coats vary in texture, color, and markings, and a specialized vocabulary has evolved to describe each characteristic.


Tail

There are many different shapes of dog tails: straight, straight up, sickle, curled and cork-screw. In some breeds, the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammal ...
is traditionally docked to avoid injuries (especially for hunting dogs). It can happen that some puppies are born with a short tail or no tail in some breeds. Dogs have a
violet gland A Rhodesian Ridgeback (sex unknown) with "stud tail": the violet gland lost hair and appears as a dark dimple The violet gland or supracaudal gland is a gland located on the upper surface of the tail of certain mammals, including European badger ...
or supracaudal gland on the dorsal (upper) surface of their tails.


Footpad

Dogs can stand, walk and run on snow and ice for long periods of time. When a dog's footpad is exposed to the cold, heat loss is prevented by an adaptation of the blood system that recirculates heat back into the body. It brings blood from the skin surface and retains warm blood in the pad surface.


Senses


Vision

Like most mammals, dogs have only two types of cone photoreceptor, making them
dichromats Dichromacy (from Greek ''di'', meaning "two" and ''chromo'', meaning "color") is the state of having two types of functioning photoreceptors, called cone cells, in the eyes. Organisms with dichromacy are called dichromats. Dichromats requir ...
. These
cone cells Cone cells, or cones, are photoreceptor cells in the retinas of vertebrate eyes including the human eye. They respond differently to light of different wavelengths, and the combination of their responses is responsible for color vision. Cone ...
are maximally sensitive between 429 nm and 555 nm. Behavioural studies have shown that the dog's visual world consists of yellows, blues and grays, but they have difficulty differentiating red and green making their color vision equivalent to red–green
color blindness Color blindness or color vision deficiency (CVD) is the decreased ability to see color or differences in color. It can impair tasks such as selecting ripe fruit, choosing clothing, and reading traffic lights. Color blindness may make some aca ...
in humans (deuteranopia). When a human perceives an object as "red," this object appears as "yellow" to the dog and the human perception of "green" appears as "white," a shade of gray. This white region (the neutral point) occurs around 480 nm, the part of the spectrum which appears blue-green to humans. For dogs, wavelengths longer than the neutral point cannot be distinguished from each other and all appear as yellow. Dogs use color instead of brightness to differentiate light or dark blue/yellow. They are less sensitive to differences in grey shades than humans and also can detect brightness at about half the accuracy of humans. The dog's visual system has evolved to aid proficient hunting. While a dog's
visual acuity Visual acuity (VA) commonly refers to the clarity of vision, but technically rates an examinee's ability to recognize small details with precision. Visual acuity is dependent on optical and neural factors, i.e. (1) the sharpness of the retinal ...
is poor (that of a
poodle The Poodle, called the Pudel in German and the Caniche in French, is a breed of water dog. The breed is divided into four varieties based on size, the Standard Poodle, Medium Poodle, Miniature Poodle and Toy Poodle, although the Medium Poodle var ...
's has been estimated to translate to a
Snellen Snellen is a Dutch surname. ''Snel'' means "quick" in Dutch and the original bearer of the name may have been a lively person. However, the origin of the surname often was patronymic, as Snel and Snelle were short forms of the archaic Germanic give ...
rating of 20/75), their visual discrimination for moving objects is very high. Dogs have been shown to be able to discriminate between humans (e.g. identifying their human guardian) at a range of between ; however, this range decreases to if the object is stationary. Dogs can detect a change in movement that exists in a single
diopter A dioptre (British spelling) or diopter (American spelling) is a unit of measurement with dimension of reciprocal length, equivalent to one reciprocal metre, 1 dioptre = 1 m−1. It is normally used to express the optical power of a lens or cur ...
of space within their eye. Humans, by comparison, require a change of between 10 and 20 diopters to detect movement.Mech, David. ''Wolves, Behavior, Ecology, and Conservation''. The University of Chicago Press, 2006, p. 98. As
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of dayli ...
hunters, dogs often rely on their vision in low light situations: They have very large pupils, a high density of rods in the
fovea Fovea () (Latin for "pit"; plural foveae ) is a term in anatomy. It refers to a pit or depression in a structure. Human anatomy *Fovea centralis of the retina * Fovea buccalis or Dimple * Fovea of the femoral head * Trochlear fovea of the fr ...
, an increased flicker rate, and a
tapetum lucidum The ''tapetum lucidum'' ( ; ; ) is a layer of tissue in the eye of many vertebrates and some other animals. Lying immediately behind the retina, it is a retroreflector. It reflects visible light back through the retina, increasing the light a ...
. The tapetum is a reflective surface behind the retina that reflects light to give the photoreceptors a second chance to catch the photons. There is also a relationship between body size and overall diameter of the eye. A range of 9.5 and 11.6  mm can be found between various breeds of dogs. This 20% variance can be substantial and is associated as an adaptation toward superior night vision. The eyes of different breeds of dogs have different shapes, dimensions, and retina configurations. Many long-nosed breeds have a "visual streak"—a wide foveal region that runs across the width of the retina and gives them a very wide field of excellent vision. Some long-muzzled breeds, in particular, the
sighthound Sighthounds, also called gazehounds, are a type of dog, hounds that hunt primarily by sight and speed, rather than by scent and endurance as scent hounds do. Appearance These dogs specialize in pursuing prey, keeping it in sight, and overp ...
s, have a field of vision up to 270° (compared to 180° for humans). Short-nosed breeds, on the other hand, have an "area centralis": a central patch with up to three times the density of nerve endings as the visual streak, giving them detailed sight much more like a human's. Some broad-headed breeds with short noses have a field of vision similar to that of humans. ] Most breeds have emmetropic, good vision, but some show a
genetic predisposition A genetic predisposition is a genetic characteristic which influences the possible phenotypic development of an individual organism within a species or population under the influence of environmental conditions. In medicine, genetic susceptibili ...
for
myopia Near-sightedness, also known as myopia and short-sightedness, is an eye disease where light focuses in front of, instead of on, the retina. As a result, distant objects appear blurry while close objects appear normal. Other symptoms may includ ...
– such as
Rottweiler The Rottweiler (, ) is a breed of domestic dog, regarded as medium-to-large or large. The dogs were known in German as , meaning Rottweil butchers' dogs, because their main use was to herd livestock and pull carts laden with butchered meat ...
s, with which one out of every two has been found to be
myopic Near-sightedness, also known as myopia and short-sightedness, is an eye disease where light focuses in front of, instead of on, the retina. As a result, distant objects appear blurry while close objects appear normal. Other symptoms may include ...
. Dogs also have a greater divergence of the eye axis than humans, enabling them to rotate their pupils farther in any direction. The divergence of the eye axis of dogs ranges from 12–25° depending on the breed. Experimentation has proven that dogs can distinguish between complex visual images such as that of a cube or a prism. Dogs also show attraction to static visual images such as the silhouette of a dog on a screen, their own reflections, or videos of dogs; however, their interest declines sharply once they are unable to make social contact with the image.


Hearing

The frequency range of dog hearing is between 16-40  Hz (compared to 20–70 Hz for humans) and up to 45–60 kHz (compared to 13–20 kHz for humans), which means that dogs can detect sounds far beyond the upper limit of the human auditory spectrum. Dogs have ear mobility that allows them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate, raise, or lower a dog's ear. A dog can identify a sound's location much faster than a human can, as well as hear sounds at four times the distance.


Smell

While the human brain is dominated by a large
visual cortex The visual cortex of the brain is the area of the cerebral cortex that processes visual information. It is located in the occipital lobe. Sensory input originating from the eyes travels through the lateral geniculate nucleus in the thalamus ...
, the dog brain is dominated by a large
olfactory cortex The olfactory system, or sense of smell, is the sensory system used for smelling (olfaction). Olfaction is one of the special senses, that have directly associated specific organs. Most mammals and reptiles have a main olfactory system and an ...
. Dogs have roughly forty times more smell-sensitive receptors than humans, ranging from about 125 million to nearly 300 million in some dog breeds, such as bloodhounds.


Taste

Dogs have around 1,700 taste buds compared to humans with around 9,000. The sweet taste buds in dogs respond to a chemical called furaneol which is found in many fruits and in tomatoes. It appears that dogs do like this flavor and it probably evolved because in a natural environment dogs frequently supplement their diet of small animals with whatever fruits happen to be available. Because of dogs' dislike of bitter tastes, various sprays, and gels have been designed to keep dogs from chewing on furniture or other objects. Dogs also have taste buds that are tuned for water, which is something they share with other carnivores but is not found in humans. This taste sense is found at the tip of the dog's tongue, which is the part of the tongue that they curl to lap water. This area responds to water at all times, but when the dog has eaten salty or sugary foods the sensitivity to the taste of water increases. It is proposed that this ability to taste water evolved as a way for the body to keep internal fluids in balance after the animal has eaten things that will either result in more urine being passed or will require more water to adequately process. It certainly appears that when these special water taste buds are active, dogs seem to get an extra pleasure out of drinking water, and will drink copious amounts of it.


Touch

The main difference between human and dog touch is the presence of specialized
whiskers Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarse ...
known as vibrissae. Vibrissae are present above the dog's eyes, below their jaw, and on their muzzle. They are sophisticated sensing organs. Vibrissae are more rigid and embedded much more deeply in the skin than other hairs and have a greater number of receptor cells at their base. They can detect air currents, subtle vibrations, and objects in the dark. They provide an early warning system for objects that might strike the face or eyes, and probably help direct food and objects towards the mouth.


Magnetic sensitivity

Dogs may prefer, when they are off the leash and Earth's magnetic field is calm, to urinate and defecate with their bodies aligned on a north–south axis. Another study suggested that dogs can see the earth's magnetic field.


Temperature regulation

Primarily, dogs regulate their body temperature through panting and sweating via their paws. Panting moves cooling air over the moist surfaces of the tongue and lungs, transferring heat to the atmosphere. Dogs and other
canid Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...
s also possess a very well-developed set of nasal turbinates, an elaborate set of bones and associated soft-tissue structures (including arteries and veins) in the
nasal cavities The nasal cavity is a large, air-filled space above and behind the nose in the middle of the face. The nasal septum divides the cavity into two cavities, also known as fossae. Each cavity is the continuation of one of the two nostrils. The nasal c ...
. These turbinates allow for heat exchange between small arteries and veins on their maxilloturbinate surfaces (the surfaces of turbinates positioned on
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...
bone) in a counter-current heat-exchange system. Dogs are capable of prolonged chases, in contrast to the ambush predation of cats, and these complex turbinates play an important role in enabling this (cats only possess a much smaller and less-developed set of nasal turbinates). This same complex turbinate structure helps conserve water in arid environments. The water conservation and thermoregulatory capabilities of these well-developed turbinates in dogs may have been crucial adaptations that allowed dogs (including both domestic dogs and their wild prehistoric ancestors) to survive in the harsh
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
environment and other cold areas of northern Eurasia and North America, which are both very dry and very cold.


References


Further reading

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External links

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