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''Malus'' ( or ) is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
of about 30–55
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of small
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, afte ...
tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondar ...
s or
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s in the family
Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are '' Alchemilla'' (270), ''Sorbus ...
, including the domesticated orchard apple, crab apples,
wild apple ''Malus sieversii'' is a wild apple native to the mountains of Central Asia in southern Kazakhstan. It has recently been shown to be the primary ancestor of most cultivars of the domesticated apple (''Malus domestica''). It was first described a ...
s, and rainberries. The genus is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entertai ...
to the
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
zone of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's North ...
.


Description

Apple trees are typically talI at maturity, with a dense, twiggy crown. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
are long, alternate, simple, with a serrated margin. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are borne in
corymb Corymb is a botanical term for an inflorescence with the flowers growing in such a fashion that the outermost are borne on longer pedicels than the inner, bringing all flowers up to a common level. A corymb has a flattish top with a superficial re ...
s, and have five
petal Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...
s, which may be white, pink, or red, and are perfect, with usually red stamens that produce copious
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophy ...
, and a half-inferior ovary; flowering occurs in the spring after 50–80
growing degree day Growing degree days (GDD), also called growing degree units (GDUs), are a heuristic tool in phenology. GDD are a measure of heat accumulation used by horticulturists, gardeners, and farmers to predict plant and animal development rates such as th ...
s (varying greatly according to subspecies and cultivar). Many apples require
cross-pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
between individuals by
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of ...
s (typically
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfami ...
s, which freely visit the flowers for both
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualis ...
and
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophy ...
); these are called self-sterile, so self-pollination is impossible, making pollinating insects essential. A number of cultivars are self-pollinating, such as 'Granny Smith' and 'Golden Delicious', but are considerably fewer in number compared to their cross-pollination dependent counterparts. Several ''Malus'' species, including domestic apples, hybridize freely. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particul ...
is a globose
pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince. Etymology The word ''pome'' entered English in the late 14th century, and ...
, varying in size from in diameter in most of the wild species, to in ''M. sylvestris sieversii'', in ''M. domestica'', and even larger in certain cultivated orchard apples. The centre of the fruit contains five
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils' ...
s arranged star-like, each containing one or two
seed A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...
s.


Subdivisions and species

About 42 to 55 species and natural hybrids are known, with about 25 from China, of which 15 are endemic. The genus ''Malus'' is subdivided into eight sections (six, with two added in 2006 and 2008). The genus ''
Docynia ''Docynia'' (栘𣐿属, ''yí yī shǔ'') is a genus of flowering trees, evergreen or semi-evergreen, in the family Rosaceae. The fruit is a pome. The tree is endemic to Southeast Asia, including Myanmar Myanmar, ; UK pronunciations: U ...
'' has been shown to be nested within ''Malus'' in molecular phylogenies. The oldest fossils of the genus date to the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...
(
Lutetian The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Midd ...
), which are leaves belonging to the species ''Malus collardii'' and ''Malus kingiensis'' from western North America (Idaho) and the
Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admin ...
(
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
), respectively.


Natural hybrids

* '' Malus × micromalus'' – midget crabapple


Fossil species

After * ''Malus collardii'' Axelrod, North America (Idaho),
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...
* ''Malus kingiensis'' Budants, Kamchatka Peninsula, Russia, Eocene * ''Malus florissantensis'' (Cockerell) MacGinitie
Green River Formation The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine ...
, North America (Colorado) Eocene * ''Malus pseudocredneria'' (Cockerell) MacGinitie Green River Formation, North America (Colorado) Eocene * ''Malus idahoensis'' R.W.Br. North America (Idaho),
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
* ''Malus parahupehensis'' J.Hsu and R.W.Chaney
Shanwang The Shanwang National Geology Park () is located in central Shandong province, People's Republic of China, about from Linqu County. It has an area of about . The Park is well known for its fossil bearing diatomitic deposits, one of only a few such ...
, Shandong, China, Miocene * ''Malus antiqua'' Doweld Romania,
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...


Cultivation

Crabapples are popular as compact ornamental trees, providing blossom in spring and colourful fruit in autumn. The fruits often persist throughout winter. Numerous hybrid cultivars have been selected. Some crabapples are used as
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
s for domestic apples to add beneficial characteristics. For example, the rootstocks of ''
Malus baccata ''Malus baccata'' is an Asian species of apple known by the common names Siberian crab apple, Siberian crab, Manchurian crab apple and Chinese crab apple. It is native to much of northern Asia, but is also grown elsewhere as an ornamental tree an ...
'' varieties are used to give additional cold hardiness to the combined plants for orchards in cold northern areas. They are also used as
pollinizer {{Unreferenced, date=May 2009 A pollenizer (or polleniser), sometimes pollinizer (or polliniser, see spelling differences) is a plant that provides pollen. The word '' pollinator'' is often used when ''pollenizer'' is more precise. A pollinato ...
s in apple
orchard An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit- or nut-producing trees which are generally grown for commercial production. Orchards are also sometimes a feature of lar ...
s. Varieties of crabapple are selected to bloom contemporaneously with the apple variety in an orchard planting, and the crabs are planted every sixth or seventh tree, or limbs of a crab tree are grafted onto some of the apple trees. In emergencies, a bucket or drum bouquet of crabapple flowering branches is placed near the beehives as orchard pollenizers. Because of the plentiful blossoms and small fruit, crabapples are popular for use in
bonsai Bonsai ( ja, 盆栽, , tray planting, ) is the Japanese art of growing and training miniature trees in pots, developed from the traditional Chinese art form of ''penjing''. Unlike ''penjing'', which utilizes traditional techniques to produce ...
culture.


Cultivars

These cultivars have won the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (N ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
:- *'Adirondack' *'Butterball' * 'Comtesse de Paris' *'Evereste' *'Jelly King'='Mattfru' *'Laura' *''
Malus × robusta ''Malus'' × ''robusta'', called Siberian crab apple along with other members of its genus, is a hybrid species of flowering plant in the family Rosaceae. It is the result of a cross between '' Malus baccata'' (Siberian crab apple) and '' Malus ...
'' 'Red Sentinel' *'Sun Rival' Other varieties are dealt with under their species names.


Toxicity

The seeds contain
cyanide Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of a ...
compounds.


Uses

Crabapple fruit is not an important
crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydroponi ...
in most areas, being extremely
sour The gustatory system or sense of taste is the sensory system that is partially responsible for the perception of taste (flavor). Taste is the perception produced or stimulated when a substance in the mouth reacts chemically with taste receptor ...
due to
malic acid Malic acid is an organic compound with the molecular formula . It is a dicarboxylic acid that is made by all living organisms, contributes to the sour taste of fruits, and is used as a food additive. Malic acid has two stereoisomeric forms (L ...
(which like the genus derives from the Latin name '' mālum''), and in some species woody, so is rarely eaten raw. In some
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
n cultures, they are valued as a sour
condiment A condiment is a preparation that is added to food, typically after cooking, to impart a specific flavor, to enhance the flavor, or to complement the dish. A table condiment or table sauce is more specifically a condiment that is served separat ...
, sometimes eaten with salt and chilli or
shrimp paste Shrimp paste or prawn sauce is a fermented condiment commonly used in Southeast Asian and Southern Chinese cuisines. It is primarily made from finely crushed shrimp or krill mixed with salt, and then fermented for several weeks. They are eith ...
. Some crabapple varieties are an exception to the reputation of being sour, and can be very sweet, such as the 'Chestnut' cultivar. Crabapples are an excellent source of
pectin Pectin ( grc, πηκτικός ': "congealed" and "curdled") is a heteropolysaccharide, a structural acid contained in the primary lamella, in the middle lamella, and in the cell walls of terrestrial plants. The principal, chemical component o ...
, and their
juice Juice is a drink made from the extraction or pressing of the natural liquid contained in fruit and vegetables. It can also refer to liquids that are flavored with concentrate or other biological food sources, such as meat or seafood, such as ...
can be made into a ruby-coloured preserve with a full, spicy flavour. A small percentage of crabapples in cider makes a more interesting flavour. As
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...
''Wergulu'', the crab apple is one of the nine plants invoked in the pagan Anglo-Saxon ''
Nine Herbs Charm The "Nine Herbs Charm" is an Old English charm recorded in the tenth-century CEGordon (1962:92–93). Anglo-Saxon medical compilation known as ''Lacnunga'', which survives on the manuscript, Harley MS 585, in the British Library, at London.Macleo ...
'', recorded in the 10th century. Applewood gives off a pleasant scent when burned, and smoke from an applewood fire gives an excellent flavour to
smoked Smoking is the process of flavoring, browning, cooking, or preserving food by exposing it to smoke from burning or smoldering material, most often wood. Meat, fish, and ''lapsang souchong'' tea are often smoked. In Europe, alder is the trad ...
foods. It is easier to cut when green; dry applewood is exceedingly difficult to carve by hand. It is a good wood for cooking fires because it burns hot and slow, without producing much flame.


Gallery

File:Apple blossom. Eastern Siberia.jpg, Apple blossom. Eastern Siberia File:Crabapples.jpg, Ripe wild crab apples (''Malus sylvestris'') File:Lee-Russell-Farm-Security-Administration-1939-Crab-Apples.jpg, Baskets of crab apples for sale in Connecticut in 1939 File:20071103Tradkrafta1.JPG, Trunk


References


External links


Germplasm Resources Information Network: ''Malus''

Flora of China: ''Malus''




a cooperative among
Purdue University Purdue University is a public land-grant research university in West Lafayette, Indiana, and the flagship campus of the Purdue University system. The university was founded in 1869 after Lafayette businessman John Purdue donated land and mon ...
,
Rutgers University Rutgers University (; RU), officially Rutgers, The State University of New Jersey, is a Public university, public land-grant research university consisting of four campuses in New Jersey. Chartered in 1766, Rutgers was originally called Queen's ...
, and the
University of Illinois The University of Illinois Urbana-Champaign (U of I, Illinois, University of Illinois, or UIUC) is a public land-grant research university in Illinois in the twin cities of Champaign and Urbana. It is the flagship institution of the Universi ...
. {{Authority control Rosaceae genera Plants used in bonsai Fruit trees Taxa named by Philip Miller