Cleaner shrimp is a
common name
In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
for a number of swimming
decapod crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s, that clean other organisms of
parasite
Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s. They belong to any of three
families
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideal ...
,
Hippolytidae
Hippolytidae is a family of cleaner shrimp, also known as broken-back shrimp or anemone shrimp. The term "broken-back shrimp" also applies to the genus '' Hippolyte'' in particular and "cleaner shrimp" is sometimes applied exclusively to ''Lysma ...
(including the
Pacific cleaner shrimp, ''Lysmata amboinensis''),
Palaemonidae (including the
spotted ''
Periclimenes magnificus''), and
Stenopodidae (including the
banded coral shrimp
''Stenopus hispidus'' is a shrimp-like decapod crustacean belonging to the infraorder Stenopodidea. Common names include coral banded shrimp and banded cleaner shrimp.
Distribution
''Stenopus hispidus'' has a pan-tropical distribution, extendi ...
, ''Stenopus hispidus'') .
The last of these families is more closely related to
lobster
Lobsters are a family (Nephropidae, synonym Homaridae) of marine crustaceans. They have long bodies with muscular tails and live in crevices or burrows on the sea floor. Three of their five pairs of legs have claws, including the first pair ...
s and
crab
Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting " tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all th ...
s than it is to the remaining families. The term "cleaner shrimp" is sometimes used more specifically for the family Hippolytidae and the genus ''
Lysmata
''Lysmata'' is a genus of shrimp in the infraorder Caridea, the caridean shrimp. The genus belongs to the family Lysmatidae. ''Lysmata'' are popular ornamental shrimp in the marine aquarium trade for their bright color patterns, interesting b ...
''.
Cleaner shrimp are so called because they exhibit a
cleaning symbiosis
Cleaning symbiosis is a mutually beneficial association between individuals of two species, where one (the cleaner) removes and eats parasites and other materials from the surface of the other (the client). Cleaning symbiosis is well-known amon ...
with client fish where the shrimp clean parasites from the fish. The fish benefit by having parasites removed from them, and the shrimp gain the nutritional value of the parasites. The shrimp also eat the mucus and parasites around the wounds of injured fish, which reduces infections and helps healing. The action of cleansing further aids the health of client fish by reducing their stress levels. In many
coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups.
...
s, cleaner shrimp congregate at
cleaning station
A cleaning station is a location where aquatic life congregate to be cleaned by smaller creatures. Such stations exist in both freshwater and marine environments, and are used by animals including fish, sea turtles and hippos, referred to as cl ...
s. In this behaviour cleaner shrimps are similar to
cleaner fish, and sometimes may join with
cleaner wrasse and other cleaner fish attending to client fish.
Shrimp of the genus ''
Urocaridella'' are often
cryptic or live in
caves
A cave or cavern is a natural void in the ground, specifically a space large enough for a human to enter. Caves often form by the weathering of rock and often extend deep underground. The word ''cave'' can refer to smaller openings such as sea ...
on the reef and are not associated
commensally with other animals.
These shrimp assemble around
cleaning station
A cleaning station is a location where aquatic life congregate to be cleaned by smaller creatures. Such stations exist in both freshwater and marine environments, and are used by animals including fish, sea turtles and hippos, referred to as cl ...
s where up to 25 shrimp live in proximity.
When a potential client fish swims close to a station with shrimp present, several shrimp perform a "rocking dance," a side-to-side movement. Frequency of rocking increases with hunger.
This increase in frequency suggests
competition
Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indiv ...
between hungry and sated shrimp.
To avoid
competition
Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indiv ...
with other cleaners during the day, the shrimp ''
Urocaridella antonbruunii'' was observed cleaning a sleeping fish at night.
Cleaner shrimps are often included in
saltwater aquaria partly due to their cleansing function and partly due to their brightly colored appearance.
References
{{reflist
Decapods
Symbiosis