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A biennial plant is a
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
that, generally in a
temperate climate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
, takes two years to complete its biological life cycle.


Life cycle

In its first year, the biennal plant undergoes primary growth, during which its vegetative structures (leaves, stems, and roots) develop. Usually, the stem of the plant remains short and the leaves are low to the ground, forming a rosette. After one year's growing season, the plant enters a period of
dormancy Dormancy is a period in an organism's life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolic activity and therefore helps an organism to conserve energy. Dormancy tends to be c ...
for the colder months. Many biennials require a cold treatment, or
vernalization Vernalization (from Latin ''vernus'', "of the spring") is the induction of a plant's flowering process by exposure to the prolonged cold of winter, or by an artificial equivalent. After vernalization, plants have acquired the ability to flower ...
, before they will flower. During the next
spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a h ...
or summer, the stem of the biennial plant elongates greatly, or "bolts". The plant then flowers, producing
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
s and
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s before it finally dies. There are far fewer biennials than either
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
s or
annual plant An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. The length of growing seasons and period in which they take place vary according to geographical ...
s. Biennials do not always follow a strict two-year life cycle and the majority of plants in the wild can take 3 or more years to fully mature. Rosette leaf size has been found to predict when a plant may enter its second stage of flowering and seed production. Alternatively, under extreme climatic conditions, a biennial plant may complete its life cycle rapidly (e.g., in three months instead of two years). This is quite common in vegetable or flower seedlings that were vernalized before they were planted in the ground. This behavior leads to many normally biennial plants being treated as annuals in some areas. Conversely, an annual grown under extremely favorable conditions may have highly successful seed propagation, giving it the appearance of being biennial or perennial. Some short-lived perennials may appear to be biennial rather than perennial. True biennials flower only once, while many perennials will flower every year once mature.


Examples

Biennials grown for flowers, fruits, or seeds are grown for two years, where those grown for edible leaves or roots are harvested after one year—and are not kept a second year to run to seed. Examples of biennial plants are members of the
onion An onion (''Allium cepa'' L., from Latin ''cepa'' meaning "onion"), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus '' Allium''. The shallot is a botanical variety of the on ...
family including
leek The leek is a vegetable, a cultivar of '' Allium ampeloprasum'', the broadleaf wild leek ( syn. ''Allium porrum''). The edible part of the plant is a bundle of leaf sheaths that is sometimes erroneously called a stem or stalk. The genus '' Al ...
, some members of the
cabbage Cabbage, comprising several cultivars of ''Brassica oleracea'', is a leafy green, red (purple), or white (pale green) biennial plant grown as an annual vegetable crop for its dense-leaved heads. It is descended from the wild cabbage ( ''B.&n ...
family, common mullein,
parsley Parsley, or garden parsley (''Petroselinum crispum'') is a species of flowering plant in the family Apiaceae that is native to the central and eastern Mediterranean region (Sardinia, Lebanon, Israel, Cyprus, Turkey, southern Italy, Greece, ...
, fennel, '' Lunaria'', silverbeet, black-eyed Susan,
sweet William Sweetness is a basic taste most commonly perceived when eating foods rich in sugars. Sweet tastes are generally regarded as pleasurable. In addition to sugars like sucrose, many other chemical compounds are sweet, including aldehydes, ketones ...
, colic weed,
carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in color, though purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild carrot, ''Daucus carota'', na ...
, and some
hollyhock ''Alcea'' is a genus of over 80 species of flowering plants in the mallow family Malvaceae, commonly known as the hollyhocks. They are native to Asia and Europe. The single species of hollyhock from the Americas, the streambank wild hollyhock, ...
s. Plant breeders have produced annual cultivars of several biennials that will flower the first year from seed, for example, foxglove and stock.


See also

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References

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